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Anthony's Keys | Family List | Rosaceae | Cotoneaster

13. Cotoneaster verruculosus Diels, Notes Roy. Bot. Gard. Edinburgh. 5: 272. 1912. Type (fr.): China. N.W. Yunnan, side valley on the eastern flank of the Tali-Range, 25°40'N., 2440–3050 m, G. Forrest 4427 (Holotype: E, isotype: A).

Cotoneaster distichus Lange var. verruculosus (Diels) T. T. Yü, Bull. Brit. Mus. (Nat. Hist.), Bot. 1: 128. 1954.

Cotoneaster sandakphuensis G. Klotz, Bull. Bot. Surv. India 5: 213. 1964; Type: "Sikkim-Himalaya, prope Sandakphu, infruticetis via, Rimbick," 9 Oct 1941, K. P. Biswas 5721, fr.

Shrubs, semi-evergreen or evergreen, prostrate to erect, (0.2–)0.6–1.2(–2) m tall; branches erect or ascending. Branchlets spiralled or distichous, initially dark reddish brown to grayish brown, dark gray when old, terete, initially densely yellow strigose, glabrate, conspicuously verruculose when old. Petiole 2–5 mm, pubescent; stipules persistent, lanceolate, pubescent; leaf blade dark green adaxially, suborbicular or broadly ovate to broadly obovate, 0.7–1.4 cm × (4–)6–12 mm, subleathery, midvein raised abaxially, impressed adaxially, both surfaces appressed pilose, glabrescent, base broadly cuneate or rounded, apex emarginate or partly mucronulate. Inflorescences usually 1-flowered. Flowers 7–8 mm in diam., nearly sessile. Hypanthium glabrous abaxially. Sepals ovate-triangular, apex ± obtuse. Petals erect, pink, suborbicular, 4–4.5 mm in diam. Stamens 20, shorter than petals; anthers white. Styles 2, not exceeding stamens. Fruit red, globose, 7–8 mm in diam.; nutlets 2(or 3). Fl. May–Jun, fr. Sep–Oct.

Mixed forests on dry slopes, grasslands, fields, roadsides; [2800–]3000–3400[–3600] m. Bhutan, W. China (Sichuan, Xizang/Tibet, Yunnan), N. and NE. India (incl. Sikkim), Myanmar, Nepal.

Specimens examined at A: Nepal: Above Dhorpatan, 3353 m, 13 Jul 1954, J. D. A. Stainton, V. R. Sykes & L. Williams 3494; East: Koshi Zone, Sankhuwa Sabha Distr., 3240 m, 27°10'N. 87°30'E., 23 Aug 1997, S. Noshiro, N. Acharya, Y. Ibaragi, K. Kobayashi & T. Kurosawa 9755196.

Further research is needed on the identity of Cotoneaster sandakphuensis and the distributional range of C. verruculosus.

stem: posture: erect, prostrate

plant: height: 20-200 cm tall

stem: branchlet: color: brown, dark, gray, red

stem: branchlet: indumentum: type: glabrate, strigose, verruculose

leaf: petiole: length: 2-5 mm

leaf: blade: shape: obovate, orbicular, ovate

leaf: blade: length: 0.7-1.4 cm long

leaf: blade: width: 0.4-1.2 cm wide

leaf: blade: length: width ratio: 1.1-1.8

leaf: texture: leathery

leaf: blade: abaxial surface: indumentum type: glabrescent, pilose

leaf: blade: adaxial surface: indumentum type: glabrescent, pilose

leaf: blade: base: cuneate, rounded

leaf: blade: apex: emarginate, mucronulate

inflorescence: type: solitary flowers

petal: color: pink

inflorescence: flowers: number: 1 (1-1)

petal: posture: erect

hypanthium: indumentum: type: glabrous

stamens: number: 20

anthers: color: white

fruit: color: red

fruit: shape: globose

fruit: diam.: 7-8 mm

nutlets: number: 2-3

flowering: May, Jun

fruiting: Sep, Oct

distribution: elevation: 2800-3600 m


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