7. Cotoneaster integrifolius (Roxburgh) G. Klotz, Wiss. Zeitschr. Mart. -Luth.Univ. Halle, Math.-Nat., xii. (Neue oder Krit.Coton.-Art.). 779. 1963.
Crataegus integrifolia Roxburgh, Fl. Ind. 2: 509. 1832; Type: India, Uttar Pradesh, Kumaon, "found by Colonel Hardwicke on the hills between Hurdwar and Sirinagur," N. Wallich 662B (left hand specimen; Herb. Roxburgh, lectotype: K).
Cotoneaster buxifolius var. marginatus Lindley ex Loudon, Encycl. Trees 411. 1842.
Cotoneaster congestus Baker in Saunders, Refug. Bot. 1: t. 51. 1869; Type: India (Sikkim), reg. Temp. & alp., 2743–4267 m, 1845, J. D. Hooker (Holotype: K).
Cotoneaster lanatus Jacques, J. Soc. Imp. Centr. Horti. Paris, t. 5: 476. 1859, nom. illeg., not Otto (1858).
Cotoneaster linearifolius (G. Klotz) G. Klotz, Wiss. Zeitschr. Friedrich-Schiller-Univ. Jena, Mat. Naturwiss. (Beitr. Phytotax., 5), 27(1): 20. 1978.
Cotoneaster marginatus (Lindley ex Loudon) Schlechtendal, Linnaea 27: 541 (1854 publ. 1856), as 'marginata', Lectotype: "C. microphylla Hort. Bot. Berold." (HAL), (selected by Klotz 1963).
Cotoneaster meuselii G. Klotz, Wiss. Zeitschr. Mart. -Luth.Univ. Halle, Math.-Nat., xii. (Neue oder Krit.Coton.-Art.) 12(10): 777. 1963; Type: "Himalaja occidentalis in Cedro-Querceto incanae sicco prope urbem Mussoori," 13 Oct 1962, H. Meusel & Schubert s.n. (Holotype: HAL).
Cotoneaster microphyllus Wallich ex Lindley var. buxifolius Dippel, Handb. Laubholzk. 3: 420. 1893.
Cotoneaster microphyllus Wallich ex Lindley var. buxifolius Dippel f. lanatus Dippel, Handb. Laubholzk. 3: 420. 1893.
Cotoneaster microphyllus Wallich ex Lindley f. linearifolius G. Klotz, Wiss. Zeitschr. Mart.-Luth. Univ. Halle, Math.-Nat. 6: 982. 1957; Type: "Planta in Horto Botanico Halensis Culta," 5 Jun 1956, G. Klotz s.n. (Holotype: JE).
Cotoneaster microphyllus Wallich ex Lindley var. thymifolius (Baker) Koehne, Mitt. Deutsch. Dendrol. Ges. 227. 1893.
Cotoneaster poluninii G. Klotz, Wiss. Zeitschr. Friedrich-Schiller-Univ. Jena, Mat. Naturwiss. (Beitr. Phytotax., 5), 27(1): 21. 1978; Type: Nepal: "Bheri River, Dunaihi, river terraces in dry locality," 2850 m, 2 Jun 1952, O. V. Polunin, W. R. Sykes & L. H. J. Williams 1037 (Holotype: A ex BM).
Cotoneaster prostratus Baker in Saunders, Refug. Bot. 1: t. 53. 1869. Type: Western Himalaya, Royle s.n. (t, 53 based on plants raised in Saunder's Garden in London from seeds sent by Royle from Western Himalaya).
Cotoneaster prostratus Baker var. lanatus (Dippel) Rehder, Man. Cult. Trees 361. 1927, as "lanata."
Cotoneaster rotundifolius Wallich ex Lindley var. lanatus (Dippel) C. K. Schneider, Ill. Handb. Laubholzk. 1: 759. 1906.
Cotoneaster thymifolius Baker in Saunders, Refug. Bot. 1: t. 50. 1869.
Shrubs, evergreen or semi-evergreen, small, (0.2–)0.5–1.5(–5) m tall, divaricate or squarrose, with regular branching; branches rigid, ± erect or decumbent. Branchlets spiralled, purple-black or green, terete, initially strigose. Leaves spiralled; petiole 1–5 mm, strigose; leaf blade shiny and pale to dark green adaxially, lanceolate to oblong or ovate, 0.4–1.7(–2.9) cm × (1.5–)3–8(–17) mm, leathery or papery, abaxially tomentose, pubescent or glabrescent, adaxially subglabrous, midvein raised abaxially, impressed adaxially, base cuneate, margin ± revolute, apex acute or acuminate, sometimes obtuse. Inflorescences 1–10-flowered or more. Pedicel 3–7 mm. Hypanthium campanulate, strigose. Sepals ovate-triangular, acute or obtuse, sometimes mucronulate. Petals spreading, white or pinkish white, suborbicular, glabrous. Stamens 20, erect; anthers reddish purple or violet-black. Styles 2(or 3). Fruit red or crimson, depressed-globose or globose, 7–9 mm; nutlets (1 or)2, rarely 3. Fl. May–Jul, fr. Oct–Nov. 2n = 51, 68.
High mountain areas; 1100–3500[–3900] m. Bhutan, W. China (Guizhou, Sichuan, S.E. Xizang/Tibet, Yunnan); India (incl. Sikkim), Kashmir, Myanmar, Nepal.
Specimens examined at A: Nepal: Pulanto, 7000 ft., 20 May 1951, O. Polunin, W. R. Sykes & L. J. H. Williams 808; West: Khaptar Forest, 3 Jun 1929, Bis Ram 110-527.
The treatment of this taxon follows Dickoré & Kasperek (In Press) who noted that this taxon has been frequently confused with other Cotoneaster taxa, particularly C. microphyllus.
plant: height: 20-500 cm tall
stem: branchlet: color: black, green, purple
stem: branchlet: indumentum: type: glabrescent, strigose
leaf: petiole: length: 1-5 mm
leaf: blade: shape: lanceolate, oblong, ovate
leaf: blade: length: 0.4-2.9 cm long
leaf: blade: width: 0.1-1.7 cm wide
leaf: blade: length: width ratio: 1.3-2.7
leaf: texture: leathery, papery
leaf: blade: abaxial surface: indumentum type: glabrous, pubescent, tomentose
leaf: blade: adaxial surface: indumentum type: glabrous
leaf: blade: base: cuneate
leaf: blade: apex: acuminate, acute, obtuse
inflorescence: type: corymb, cyme, fascicled flowers, solitary flowers
petal: color: pink, white
inflorescence: flowers: number: 1-10
petal: posture: spreading
pedicel: length: 3-7 mm
hypanthium: indumentum: type: strigose
stamens: number: 20 (20-20)
anthers: color: black, purple, red, violet
fruit: color: crimson, red
fruit: shape: depressed, globose
fruit: diam.: 7-9 mm
nutlets: number: 1-3
flowering: May, Jun, Jul
fruiting: Oct, Nov
distribution: elevation: 1100-3900 m