Distribution: Endemic to Madagascar

Bioclimate: Humid, Subhumid

Vegetation Formation: Forest

Elevation (m): 0-499, 500-999

Protected Areas : Analamazaotra-Périnet, Anjanaharibe-Sud, Betampona, Mantadia, Marojejy, Masoala, Zahamena

Note: We provisionally accept a broad concept for this sp, but as such is very variable, and the variation certainly shows some ecogeographic patterns. Further work may result in splitting the sp into 2 or more separate taxa

Updated on: 10 Oct. 2008