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FOC | Family List | FOC Vol. 7 | Lauraceae

14. Machilus Rumphius ex Nees in Wallich, Pl. Asiat. Rar. 2: 61, 70. 1831.

润楠属 run nan shu

Authors: Fa-Nan Wei & Henk van der Werff

Evergreen trees or shrubs. Leaves alternate, entire, pinnately veined. Inflorescences usually paniculate, terminal, subterminal, or arising from near base of branchlet, with long peduncles or rarely without peduncles. Flowers bisexual, small or larger. Perianth tubes short; perianth lobes 6 in 2 series, equal, subequal, or occasionally outer ones conspicuously smaller than inner ones, usually persistent, rarely deciduous. Fertile stamens 9 in 3 series, anthers 4-celled, 1st and 2nd series of stamens eglandular, anthers introrse, 3rd series of stamens glandular, anthers extrorse or lateral, glands stipitate to sessile. Staminodes in 4th series, sagittate. Ovary sessile; stigmas small, dish-shaped or capitate. Fruit fleshy, globose, rarely ellipsoid or oblong, subtended at base by persistent and reflexed perianth lobes; fruiting pedicel not enlarged or slightly enlarged.

About 100 species: tropical and subtropical S and SE Asia; 82 species (63 endemic) in China; one additional species (endemic) is of uncertain placement.

The following species were described from China but could not be treated here because no material was seen by the present authors:

Machilus hemsleyi Nakai (J. Jap. Bot. 16: 122. 1940; Persea hemsleyi (Nakai) Kostermans), described from Hubei.

Machilus tingzhourensis M. M. Lin et al. (Bull. Bot. Res., Harbin 25: 5. 2005), described from Fujian.

Species incertae sedis

Machilus holadena H. Liu, Laurac. Chine & Indochine, 54. 1932 ["holadenus"].
全腺润楠 quan xian run nan

Trees. Branchlets purple-brown, slender, ± angular, subglabrous. Terminal bud small; bud scales 3-5 mm. Leaves subopposite; petiole 1-1.5 cm; leaf blade dull abaxially, shiny adaxially, elliptic, 10-11 × 3.5-4.5 cm, leathery, glabrous on both surfaces, midrib and lateral veins conspicuous on both surfaces, lateral veins ca. 10 pairs, veinlets anastomosing, somewhat visible abaxially, base cuneate, apex acuminate, acumen ca. 1 cm. Cymose panicles few flowered, inserted on short shoots in upper leaf axils or in axils of deciduous bracts; short shoots 2.5-5 mm; peduncles 1.5-4.5 cm. Pedicel 5-10 mm (terminal one longer), glabrous. Flowers yellow. Perianth tube long; lobes (6-)9 or 12, oblong, equal, 3.5-5 mm, glabrous abaxially, pilose adaxially, glandular. Stamens 9 or 12, 3-4 mm, all 2-glandular; glands at middle part of filament, stipitate, sublanceolate; filaments pilose; anthers subovoid, 2(or 4)-celled. Staminodes 3, somewhat pilose, apex deltoid. Pistil 4.5-5 mm, glabrous; ovary subovoid, almost covered by perianth tube, apex attenuate into a style 2.5-3 mm; stigma 3-lobed. Fruit unknown. Fl. May.

● Mountain slopes. Sichuan (Emei Shan), NE Yunnan.

Li Jie and Li Xiwen note that this species has subopposite leaves, a long perianth tube, (6-)9 or 12 perianth lobes, and mostly 2-celled (rarely 4-celled) anthers. They consider that it may perhaps be a species of Beilschmiedia, but they have not yet seen its fruit and therefore include it here as a species incertae sedis.

Key 6

1 Fruit ovoid to suboblong   (2)
+ Fruit globose or compressed globose   (3)
2 (1) Fruit ovoid; leaf blade elliptic, 8-13 × 2.5-5 cm, lateral veins 8-10 pairs.   71 M. melanophylla
+ Fruit suboblong; leaf blade obovate, oblong, or oblanceolate, 13-24 × 5.5-8(-10) cm, lateral veins 14-16 pairs.   72 M. bonii
3 (1) Perianth lobes unequal   (4)
+ Perianth lobes equal or subequal   (7)
4 (3) Leaf blade puberulent abaxially when mature, densely golden yellow sericeous on both surfaces when young, narrowly lanceolate, 7-17 × 1.8-3.5 cm.   73 M. viridis
+ Leaf blade glabrous abaxially when mature, glabrous at least adaxially when young, wider, 5-6 cm wide   (5)
5 (4) Flowers large, to 9 mm; leaf blade lateral veins dense, 15-20 pairs.   74 M. rufipes
+ Flowers smaller; leaf blade lateral veins more sparse, 5-12 pairs   (6)
6 (5) Perianth lobes oblong-elliptic, inner ones ca. 5.5 mm; leaf blade elliptic to oblanceolate or elliptic-lanceolate, 11-16 × 2-4 cm, lateral veins 10-12 pairs.   75 M. oculodracontis
+ Perianth lobes ovate, inner ones ca. 3.5 mm; leaf blade obovate or obovate-elliptic to elliptic, 6-12.5 × 2-5.6 cm, lateral veins 5-7 pairs.   76 M. monticola
7 (3) Leaf blade abaxially densely gray-brown tomentose; fruit compressed globose.   77 M. velutinoides
+ Leaf blade glabrous on both surfaces or appressed puberulent abaxially; fruit globose   (8)
8 (7) Involucral bracts persistent in flowering stage; leaf blade lateral veins 16-20 pairs.   78 M. shweliensis
+ Involucral bracts deciduous in flowering stage; leaf blade lateral veins (5-)9-13 pairs   (9)
9 (8) Leaf apex obtuse or rotund.   79 M. pomifera
+ Leaf apex subacute to acuminate   (10)
10 (9) Petiole 2.5-5 cm, 2-3.5 mm thick; leaf blade elliptic to obovate-elliptic or suboblong; flowers large, 7-8(-10) mm.   80 M. robusta
+ Petiole 1-2 cm, more slender; leaf blade elliptic to oblong; flowers smaller, 2.5-3.5 mm   (11)
11 (10) Leaf blade 6-15.5 × 2-4.8 cm; panicles 2.5-5.5 cm, puberulent; perianth lobes puberulent on both surfaces.   81 M. verruculosa
+ Leaf blade 10.5-20 × 3.5-7 cm; panicles 12-15 cm, subglabrous; perianth lobes glabrous outside.   82 M. ovatiloba

List of Keys

  • List of lower taxa


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