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FNA | Family List | FNA Vol. 2 | Schizaeaceae

2. Actinostachys Wallich, Numer. List. 1. 1829.

Ray spiked fern [Greek aktis, ray, and stachys, spike, referring to the rays of the fertile leaves]

Warren H. Wagner Jr.

Plants terrestrial. Roots dark, fibrous, covered with dark, stiff hairs, 2--3 mm. Stems upright; hairs uniseriate. Leaves all fertile (even youngest), unbranched, long-petioled. Blades falsely digitate, reduced to 2--many erect to spreading terminal rays; rays appearing to be whorled but actually borne on very short rachis. Sporangia in 2--4 rows. Gametophytes subterranean, not green, tuberlike, brown-hairy. x = 134, 140.

Species 20 (1 in the flora): nearly worldwide in tropical regions.


Wagner, W. H. Jr. and V. Quevedo. 1985. Polymorphism in Actinostachys pennula (Swartz) Hooker and the taxonomic status of A. germanii (Fée) Prantl. [Abstract.] Amer. J. Bot. 72: 927--928.

Lower Taxon


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