17. Clethraceae Klotzsch
Gordon C. Tucker, Sean C. Jones
Shrubs [trees]. Leaves deciduous [persistent], cauline, alternate, simple; stipules absent; petiole present; blade margins serrate. Inflorescences terminal racemes, solitary or in clusters. Flowers bisexual; perianth and androecium hypogynous; sepals 5, connate proximally; petals 5, connate proximally ca. 1/2 their lengths, corolla cylindric-urceolate; nectary disc absent; stamens 10, in 2 whorls of 5, outer whorl antipetalous, free, distinct or barely adnate to petals; anthers dehiscent by apical porelike slits; pistils 1, 3-carpellate; ovary superior, 3-locular; placentation axile; ovules anatropous, unitegmic, tenuinucellate; styles 1, hollow; stigmas 3, linear. Fruits capsular, dehiscence loculicidal, enclosed by persistent calyx. Seeds [6-]40-100, brown, oblong-ovoid; embryo straight; endosperm oily, well differentiated.
Genera 2, species ca. 75 (1 genus, 3 species in the flora): e North America, Mexico, Central America, South America, e Asia, Atlantic Islands (Madeira).
Clethraceae is often treated to include only Clethra; studies have shown that Purdiaea Planchet, a neotropical genus of about 12 species (most diverse in Cuba), should be included in Clethraceae rather than Cyrillaceae (J. V. Schneider and C. Bayer 2004). This is consistent with the molecular interpretation of Clethraceae as a sister group to Cyrillaceae and Ericaceae.
SELECTED REFERENCES Schneider, J. V. and C. Bayer. 2004. Clethraceae. In: K. Kubitzki et al., eds. 1990+. The Families and Genera of Vascular Plants. 9+ vols. Berlin etc. Vol. 6, pp. 69-73. Thomas, J. L. 1961. The genera of the Cyrillaceae and Clethraceae of the southeastern United States. J. Arnold Arbor. 42: 96-106.