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143. Eucephalus Nuttall, Trans. Amer. Philos. Soc., n. s.  7: 298.  1840.  
Aster [Greek eu-, good or original, and kephalotos, with a head; alluding "to the elegant qualities of the calyx"—T. Nuttall 1840]  
 
Geraldine A. Allen
 
 
 
Aster Linnaeus sect. Eucephalus (Nuttall) Munz & D. D. Keck ex A. G. Jones; Aster subsect. Eucephalus (Nuttall) Bentham 
Perennials, 10–160 cm (usually cespitose, induments usually of stipitate-glandular and smooth-surfaced, curved or twisted woolly hairs, plants with caudices or short rhizomes, roots fibrous). Stems ascending or erect, simple, glabrate, puberulent, pilose, cottony, or woolly, eglandular or glandular. Leaves cauline; alternate; sessile (proximal withering by flowering; proximalmost reduced, scalelike) ; blades (1-nerved) ovate, elliptic, oblong, lanceolate, or linear (± uniform in size), margins entire, faces glabrate, scabrous, cottony, or woolly, eglandular or stipitate-glandular. Heads radiate or discoid, usually in open, racemiform, paniculiform, or corymbiform arrays, sometimes borne singly. Involucres turbinate-cylindric, turbinate, turbinate-obconic, or campanulate, 10–25 mm diam. Phyllaries 20–50 in 3–6 series, ± unequal (± appressed, often reddish or purplish at margins and tips), 1-nerved (keeled), ovate, lance-oblong, lanceolate, linear-oblong, or linear, chartaceous at bases, margins sometimes hyaline, especially proximally; apices acute to obtuse, green, usually puberulent, tomentose, and/or stipitate-glandular, sometimes glabrous. Receptacles ± flat, pitted, epaleate. Ray florets 0–21 (usually 5, 8, or 13), pistillate, fertile; corollas violet-purple, purple, pink, or white . Disc florets 10–35, bisexual, fertile; corollas yellow, ± ampliate, tubes shorter than funnelform throats, lobes 5, erect or reflexed, triangular; style-branch appendages lanceolate. Cypselae ± obconic, flattened, laterally 1–2-ribbed, sometimes with 1–2 additional nerves on each face, glabrous, pilose, or strigose, eglandular; pappi persistent, of 30–50 whitish to tawny, barbellate or smooth, apically clavate or more conspicuously barbellate bristles in 2(–3) series (outer usually 1 mm or less, sometimes 0, inner 5–10 mm). x = 9. 
Species 10 (10 in the flora): North America. 
 
 
 
Eucephalus, a relatively well-marked western North American group, has been treated as a section of Aster or as a distinct genus. Recent molecular evidence places Eucephalus, together with the eastern North American Doellingeria, at the base of the North American clade of Astereae. 
 
 
 
 
                        
                         
		
                          
                          
			
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  | 1 | 
  Ray florets usually 1–4, often 0 | 
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  (2) | 
 
  | + | 
  Ray florets commonly 5, 8, or 13+ | 
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  (5) | 
 
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  | 2 (1) | 
  Ray florets 0; leaves 5–9 cm, ± glabrous abaxially, glandular adaxially; plants 60–120 cm; open woods, Lane County, Oregon | 
    | 
  10 Eucephalus vialis | 
 
  | + | 
  Ray florets usually 1–4; leaves 2–6 cm, hairy; plants 10–100 cm | 
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  (3) | 
 
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  | 3 (2) | 
  Leaves abaxially moderately to densely hairy, adaxially glabrous or nearly so | 
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  9 Eucephalus tomentellus | 
 
  | + | 
  Leaves glabrous, eglandular or sparsely glandular on both faces | 
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  (4) | 
 
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  | 4 (3) | 
  Phyllaries subequal | 
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  1 Eucephalus breweri | 
 
  | + | 
  Phyllaries strongly unequal | 
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  4 Eucephalus glabratus | 
 
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  | 5 (1) | 
  Stems, leaves, and phyllaries glabrous, glaucous; plants 40–160 cm; leaves linear tonarrowly lance-elliptic, 4–10 cm; rays purple | 
    | 
  5 Eucephalus glaucescens | 
 
  | + | 
  Stems, leaves, and phyllaries pubescent or glabrate, glandular or not, not glaucous; plants 10–120(–150) cm; leaves elliptic, oblong, lance-ovate, lance-elliptic, lanceolate, linear-oblong or -lanceolate, 1.5–10 cm; rays white, pink, violet, or purple. | 
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  (6) | 
 
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  | 6 (5) | 
  Leaves 5–10 cm, elliptic to lanceolate, glabrous and eglandular, or abaxially ± glandular and/or villous; plants 50–150 cm; rays white to pink | 
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  3 Eucephalus engelmannii | 
 
  | + | 
  Leaves 1.5–7 cm, elliptic, elliptic-oblong, oblong, lance-ovate, lance-elliptic, linear-oblong or -lanceolate, glandular or not, scabrous or cottony; plants 10–80 cm; rays white or violet to purple | 
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  (7) | 
 
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  | 7 (6) | 
  Rays white; stems pilose or sparsely to moderately glandular-pubescent | 
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  (8) | 
 
  | + | 
  Rays violet to purple; stems scabrous (to scabrellous) or cottony and/or glandular-pubescent (especially peduncles) | 
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  (9) | 
 
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  | 8 (7) | 
  Phyllaries lance-ovate; Cascade Mountains, Oregon | 
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  6 Eucephalus gormanii | 
 
  | + | 
  Phyllaries lance-linear; Olympic Mountains, Washington | 
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  8 Eucephalus paucicapitatus | 
 
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  | 9 (7) | 
  Leaves moderately scabrellous (and sometimes glandular) on both faces | 
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  2 Eucephalus elegans | 
 
  | + | 
  Leaves abaxially strongly cottony, adaxially glabrous or sparsely scabrous | 
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  7 Eucephalus ledophyllus | 
 
 
 
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