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143. Eucephalus Nuttall, Trans. Amer. Philos. Soc., n. s. 7: 298. 1840.
Aster [Greek eu-, good or original, and kephalotos, with a head; alluding "to the elegant qualities of the calyx"—T. Nuttall 1840]
Geraldine A. Allen
Aster Linnaeus sect. Eucephalus (Nuttall) Munz & D. D. Keck ex A. G. Jones; Aster subsect. Eucephalus (Nuttall) Bentham
Perennials, 10–160 cm (usually cespitose, induments usually of stipitate-glandular and smooth-surfaced, curved or twisted woolly hairs, plants with caudices or short rhizomes, roots fibrous). Stems ascending or erect, simple, glabrate, puberulent, pilose, cottony, or woolly, eglandular or glandular. Leaves cauline; alternate; sessile (proximal withering by flowering; proximalmost reduced, scalelike) ; blades (1-nerved) ovate, elliptic, oblong, lanceolate, or linear (± uniform in size), margins entire, faces glabrate, scabrous, cottony, or woolly, eglandular or stipitate-glandular. Heads radiate or discoid, usually in open, racemiform, paniculiform, or corymbiform arrays, sometimes borne singly. Involucres turbinate-cylindric, turbinate, turbinate-obconic, or campanulate, 10–25 mm diam. Phyllaries 20–50 in 3–6 series, ± unequal (± appressed, often reddish or purplish at margins and tips), 1-nerved (keeled), ovate, lance-oblong, lanceolate, linear-oblong, or linear, chartaceous at bases, margins sometimes hyaline, especially proximally; apices acute to obtuse, green, usually puberulent, tomentose, and/or stipitate-glandular, sometimes glabrous. Receptacles ± flat, pitted, epaleate. Ray florets 0–21 (usually 5, 8, or 13), pistillate, fertile; corollas violet-purple, purple, pink, or white . Disc florets 10–35, bisexual, fertile; corollas yellow, ± ampliate, tubes shorter than funnelform throats, lobes 5, erect or reflexed, triangular; style-branch appendages lanceolate. Cypselae ± obconic, flattened, laterally 1–2-ribbed, sometimes with 1–2 additional nerves on each face, glabrous, pilose, or strigose, eglandular; pappi persistent, of 30–50 whitish to tawny, barbellate or smooth, apically clavate or more conspicuously barbellate bristles in 2(–3) series (outer usually 1 mm or less, sometimes 0, inner 5–10 mm). x = 9.
Species 10 (10 in the flora): North America.
Eucephalus, a relatively well-marked western North American group, has been treated as a section of Aster or as a distinct genus. Recent molecular evidence places Eucephalus, together with the eastern North American Doellingeria, at the base of the North American clade of Astereae.
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1 |
Ray florets usually 1–4, often 0 |
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(2) |
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Ray florets commonly 5, 8, or 13+ |
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(5) |
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2 (1) |
Ray florets 0; leaves 5–9 cm, ± glabrous abaxially, glandular adaxially; plants 60–120 cm; open woods, Lane County, Oregon |
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10 Eucephalus vialis |
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Ray florets usually 1–4; leaves 2–6 cm, hairy; plants 10–100 cm |
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(3) |
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3 (2) |
Leaves abaxially moderately to densely hairy, adaxially glabrous or nearly so |
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9 Eucephalus tomentellus |
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Leaves glabrous, eglandular or sparsely glandular on both faces |
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(4) |
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4 (3) |
Phyllaries subequal |
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1 Eucephalus breweri |
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Phyllaries strongly unequal |
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4 Eucephalus glabratus |
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5 (1) |
Stems, leaves, and phyllaries glabrous, glaucous; plants 40–160 cm; leaves linear tonarrowly lance-elliptic, 4–10 cm; rays purple |
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5 Eucephalus glaucescens |
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Stems, leaves, and phyllaries pubescent or glabrate, glandular or not, not glaucous; plants 10–120(–150) cm; leaves elliptic, oblong, lance-ovate, lance-elliptic, lanceolate, linear-oblong or -lanceolate, 1.5–10 cm; rays white, pink, violet, or purple. |
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(6) |
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6 (5) |
Leaves 5–10 cm, elliptic to lanceolate, glabrous and eglandular, or abaxially ± glandular and/or villous; plants 50–150 cm; rays white to pink |
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3 Eucephalus engelmannii |
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Leaves 1.5–7 cm, elliptic, elliptic-oblong, oblong, lance-ovate, lance-elliptic, linear-oblong or -lanceolate, glandular or not, scabrous or cottony; plants 10–80 cm; rays white or violet to purple |
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(7) |
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7 (6) |
Rays white; stems pilose or sparsely to moderately glandular-pubescent |
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(8) |
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Rays violet to purple; stems scabrous (to scabrellous) or cottony and/or glandular-pubescent (especially peduncles) |
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(9) |
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8 (7) |
Phyllaries lance-ovate; Cascade Mountains, Oregon |
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6 Eucephalus gormanii |
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Phyllaries lance-linear; Olympic Mountains, Washington |
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8 Eucephalus paucicapitatus |
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9 (7) |
Leaves moderately scabrellous (and sometimes glandular) on both faces |
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2 Eucephalus elegans |
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Leaves abaxially strongly cottony, adaxially glabrous or sparsely scabrous |
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7 Eucephalus ledophyllus |
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List of lower taxa
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