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FNA | Family List | FNA Vol. 21 | Asteraceae

377. Hulsea Torrey & A. Gray in War Department [U.S.], Pacif. Railr. Rep. 6(3): 77, plate 13. 1858.

Alpinegold [For Gilbert White Hulse, 1807–1883, physician and plant collector]

Dieter H. Wilken

Annuals or perennials, 5–150 cm. Stems erect, simple or branched (aerial shoots often from subterranean caudices). Leaves basal or basal and cauline, proximally whorled, distally alternate; petiolate (at least basal) or sessile; blades mostly lance-linear, oblanceolate, ovate, or spatulate, margins entire, lobed, or toothed, faces thinly lanate to densely woolly (hairs crisped, tangled, or matted, usually 0.8+ mm) and/or gland-dotted, glandular-puberulent, glandular-villous, or stipitate-glandular. Heads radiate, borne singly or in corymbiform arrays. Involucres obconic to hemispheric, 8–26 mm diam. Phyllaries persistent, 14–35(–60+) in 2–4 series (reflexed in fruit, mostly lance-ovate to lanceolate or linear, herbaceous). Receptacles flat, knobby or pitted, epaleate. Ray florets 9–60+, pistillate, fertile; corollas yellow, orange, or red. Disc florets 20–200+, bisexual, fertile; corollas yellow to orange (glabrous or sparsely hairy), tubes shorter than or about equaling cylindric throats, lobes 5, deltate to lance-ovate. Cypselae clavate to linear, compressed (lenticular in cross section), silky-hairy; pappi of 4 (distinct) quadrate to spatulate, equal, subequal, or unequal, erose to laciniate scales (the alternate alike). x = 19.

Species 7 (7 in the flora): w United States, nw Mexico.


Wilken, D. H. 1975. A systematic study of Hulsea (Asteraceae). Brittonia 27: 228–244.

1 Basal and proximal cauline leaves gray to grayish green, lanate to woolly (hairs mostly eglandular)   (2)
+ Basal and proximal cauline leaves ± green, mostly glandular-puberulent or glandular villous (sometimes sparsely lanate as well, sometimes woolly in H. nana)   (3)
2 (1) Leaves basal and cauline, blades broadly oblanceolate to spatulate, 6–10 cm; heads 2–5; cypselae 4–6 mm   3 Hulsea californica
+ Leaves mostly basal, blades spatulate, 1–9 cm; heads 1–2; cypselae 5–10 mm   7 Hulsea vestita
3 (1) Leaves mostly basal (cauline none or relatively few); heads 1–2 (per basal leaf rosette)   (4)
+ Leaves mostly cauline (basal usually present as well); heads 3–5   (5)
4 (3) Plants (10–)20–40 cm; leaf margins sinuate, lobed, or toothed (lobes or teeth mostly triangular); phyllary apices acute to attenuate; ray florets 28–59   1 Hulsea algida
+ Plants 5–15(–20) cm; leaf margins lobed (lobes mostly oblong); phyllary apices acuminate to acute; ray florets 12–30   6 Hulsea nana
5 (3) Ray laminae red to reddish purple (narrowly oblong to linear, ciliate)   4 Hulsea heterochroma
+ Ray laminae yellow (narrowly elliptic to narrowly lance-oblong)   (6)
6 (5) Ray florets 10–23, corolla tubes hairy   2 Hulsea brevifolia
+ Ray florets 20–35, corolla tubes glabrous   5 Hulsea mexicana

  • List of lower taxa


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