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FNA | Family List | FNA Vol. 27 | Scouleriaceae

1. Scouleria Hooker, Bot. Misc. 1: 33. 1829.

[For John Scouler, 1804-1871, physician, botanical collector, and naturalist]

Grimmia sect. Scouleria (Hooker) Müller Hal.; Grimmia subgen. Scouleria (Hooker) Lesquereux & James

Plants clustered in loose to compact tufts, black or brown with age. Stems ca. 4-10 cm, central strand absent; rhizoids forming a “hold-fast,” occasionally extending from stem and abaxial surface of leaves. Leaves erect to patent when dry, when present pseudocosta marginal [submarginal]; apex often cucullate. Outer perichaetial leaves one-third shorter than inner leaves. Vaginula forming a sheath around proximal third or fourth of seta. Seta 1/3-1/2 capsule length. Capsule glossy black with age, columella when dry appearing longitudinally ribbed, when wet expanding, becoming stoutly cylindrical or ovoid. Spores 30-55 µm.

Species 3 (2 in the flora): nw North America, temperate South America, n Asia.

Scouleria patagonica) is the only taxon found in South America.

1 Leaf margin 1-stratose or partially (rarely fully) 2-stratose; peristome present.   1 Scouleria aquatica
+ Leaf margin multistratose; peristome absent.   2 Scouleria marginata

Lower Taxa


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