18. Suksdorfia A. Gray, Proc. Amer. Acad. Arts. 15: 41. 1879.
[For Wilhelm Nikolaus Suksdorf, 1850-1932, German botanist and collector in the Pacific Northwest]
Elizabeth Fortson Wells, Patrick E. Elvander
Hemieva Rafinesque, name rejected
Herbs, not rhizomatous, not stoloniferous; base of slender caudex bearing bulbils. Flowering stems erect, leafy, 8-40 cm, stipitate-glandular. Leaves in basal rosette and cauline; cauline leaves reduced distally; stipules of basal leaves inconspicuous, stipules of cauline leaves conspicuous; petiole sparsely to moderately stipitate-glandular; blade reniform to orbiculate, usually 3-9-lobed, base cordate to truncate, ultimate margins irregularly crenate, ciliate, apex obtuse to rounded, surfaces glabrous or sparsely stipitate-glandular; venation palmate. Inflorescences flat-topped panicles of simple or compound cymes, terminal from terminal bud in rosette, 2-35-flowered, bracteate or ebracteate. Flowers: hypanthium adnate to ovary in proximal 7/8, free from ovary 0.3-1 mm, greenish (with purple band distally in S. ranunculifolia); sepals 5, greenish (with purple margins in S. violacea); petals 5, white or pink, purple, or violet; nectary tissue partially covering ovary (in S. ranunculifolia) or absent (in S. violacea); stamens 5; filaments filiform; ovary 1/2-3/4 inferior, 2-locular, carpels connate proximally; placentation axile; styles absent or 2; stigmas 2. Capsules 2(-3)-beaked. Seeds (50-75), dark brown, ellipsoid and prismatic (or angular), warty. x = 7.
Species 3 (2 in the flora): w North America, South America (Argentina, Bolivia).
Suksdorfia has been split into three monospecific genera by different authors. The work of R. J. Gornall and B. A. Bohm (1980, 1984, 1985) emphasized the similarities of the species in supporting a single genus concept. More recent, molecular data (D. E. Soltis et al. 1993; L. A. Johnson and Soltis 1994) suggest that S. violacea is more closely related to species of Bolandra, that S. ranunculifolia is more closely related to species of Boykinia, and that the two North American species indeed should be placed into monospecific genera. The South American species is S. alchemilloides (Grisebach) Engler of northern Argentina and Bolivia.
