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FNA | Family List | FNA Vol. 7 | Brassicaceae

54. Tropidocarpum Hooker, Icon. Pl. 1: plate 43. 1836.

[Greek tropis, keel, and karpos, fruit, alluding to fruit shape]

Ihsan A. Al-Shehbaz

Agallis Philippi; Twisselmannia Al-Shehbaz

Annuals; not scapose; pubescent or glabrous, trichomes simple, these mixed with smaller, short-stalked, Y-shaped, forked ones, rarely rays branched. Stems (usually several from base, rarely simple), usually ascending, decumbent, or prostrate, rarely erect, unbranched or branched [basally] distally, (slender). Leaves basal and cauline; petiolate or sessile; basal (withered by fruiting), not rosulate, petiolate, blade margins pinnatifid or pinnatisect; cauline sessile or subsessile, blade margins pinnatifid. Racemes (several-flowered, bracteate throughout), considerably elongated in fruit. Fruiting pedicels erect, ascending, or divaricate, slender. Flowers: sepals erect to ascending, oblong, (glabrous or pubescent); petals yellow, sometimes with purple tinge, oblanceolate, obovate, or spatulate, (slightly longer than sepals), claw undifferentiated from blade; stamens slightly tetradynamous; filaments sometimes slightly dilated basally; anthers ovate, (apex obtuse); nectar glands confluent, subtending bases of stamens. Fruits siliques or silicles, sessile, linear, oblong, or obdeltoid, smooth, strongly angustiseptate; valves [(2 or) 4 in T. capparideum, keeled], not veined, (thin-leathery throughout, or distally subwoody or thick-leathery and tuberculate-rugose), pubescent; replum rounded; septum complete or absent; ovules 4-70 per ovary; style distinct, (slender); stigma capitate. Seeds uniseriate or (in 4-valved fruits) 4-seriate, not winged, oblong; seed coat (reticulate), mucilaginous when wetted; cotyledons incumbent. x = 8.

Species 4 (3 in the flora): California, c Mexico, South America (c Chile).

SELECTED REFERENCES Al-Shehbaz, I. A. 2003c. A synopsis of Tropidocarpum (Brassicaceae). Novon 13: 392-395. Al-Shehbaz, I. A. and R. A. Price. 2001. The Chilean Agallis and Californian Tropidocarpum (Brassicaceae) are congeneric. Novon 11: 292-293. Robinson, B. L. 1896. The fruit of Tropidocarpum. Erythea 4: 108-119.

1 Fruits obdeltoid, valves thick-leathery and tuberculate-rugose distally, puberulent, trichomes antrorse; ovules 4-8 per ovary.   3 Tropidocarpum californicum
+ Fruits linear or oblong, valves thin-leathery and smooth distally, usually pubescent or puberulent, rarely glabrescent, trichomes retrorse; ovules 25-70 per ovary   (2)
2 (1) Fruits narrowly linear, (25-)30-60(-70) × 1.5-2(-3) mm, length 13-46 times width; valves 2.   1 Tropidocarpum gracile
+ Fruits oblong, (5-)9-20 × (3-)4-5 mm, length (1.6-)2.8-5 times width; valves (2 or) 4.   2 Tropidocarpum capparideum

Lower Taxa


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