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FNA | Family List | FNA Vol. 28 | Leskeaceae | Pseudoleskea

5. Pseudoleskea radicosa (Mitten) Macoun & Kindberg, Cat. Canad. Pl., Musci. 181. 1892.

Hypnum radicosum Mitten, J. Linn. Soc., Bot. 8: 31. 1864; Lescuraea radicosa (Mitten) Mönkemeyer

Plants medium-sized to large, in thick mats, green, yellow-green, or orange-green. Stems with branches robust, julaceous, apices usually curving up; central strand present; paraphyllia many, filamentous to foliose, branched. Leaves appressed to somewhat erect when dry, erect-spreading when moist, glossy or dull, ovate to ovate-lanceolate, asymmetric, usually falcate to falcate-secund, 0.6-2(-2.4) mm; margins narrowly recurved to mid leaf or to near acumen; apex abruptly acute to short- or long-acuminate, hair-point absent; costa subpercurrent to percurrent, yellow-green, sometimes sinuate; alar cells transversely elongate to quadrate, region small to medium-sized; medial laminal cells , short-rhomboidal, elliptic, or fusiform, to 40 µm, 2-3(-4):1, pellucid to opaque, prorate to near base, , walls thin or rarely firm, not pitted; ; juxtacostal cells somewhat shorter than more distal cells, walls not pitted. Capsule erect to suberect, symmetric, 1-2 mm; endostome basal membrane 1/4 -1/3 exostome length, segments shorter than exostome, cilia usually present, or 2, sometimes reduced. Spores 16-22 µm.

Varieties 3 (3 in the flora): North America, Eurasia, Atlantic Islands (Iceland).

Pseudoleskea radicosa is a common and variable species found at moderate to high elevations; the capsules mature in summer. Two of the three varieties are quite distinctive. Pseudoleskea radicosa is closely related to P. incurvata but differs in the longer, more homogeneous, thin-walled, and wider laminal cells. Combined ecological, molecular and morphological studies of the varieties are needed to determine their distinctiveness.

1 Plants large; stem apices hooked; branches few or absent; leaves 1.6-2.3 mm, strongly falcate-secund, pluriplicate; apices long-acuminate.   5c Pseudoleskea radicosa var. denudata
+ Plants small to medium-sized; stem apices not distinctly hooked; branches many; leaves usually less than 1.6 mm, straight, falcate, or falcate-secund, 2-plicate; apices predominantly short-acuminate to acute   (2)
2 (1) Plants medium-sized; leaves weakly to moderately concave, not incurved, 1.2-1.7 mm; apices usually long-acuminate; medial laminal cells opaque.   5a Pseudoleskea radicosa var. radicosa
+ Plants small; leaves strongly concave, often incurved, 0.8-1.3 mm; apices acute to short-acuminate; medial laminal cells pellucid.   5b Pseudoleskea radicosa var. compacta


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