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FNA | Family List | FNA Vol. 28 | Hypnaceae

17. Pylaisia Schimper in P. Bruch and W. P. Schimper, Bryol. Europ. 5: 87, plate 455. 1851. (as Pylaisaea), name and orthography conserved • [For Auguste Jean Marie Bachelot de la Pylaie, 1786-1856, French bryologist].

Tomotsugu Arikawa

Pylaisiella Kindberg ex Grout

Plants small, in low mats, yellowish to whitish, shiny. Stems creeping, regularly or irregularly pinnate, ; hyalodermis absent, central strand present, small; pseudoparaphyllia foliose. Stem and branch leaves similar. Stem leaves appressed when dry, erect-spreading when moist, ovate-lanceolate, , not or slightly plicate; base not or weakly decurrent; margins plane, sometimes slightly involute distally, subentire to serrulate distally; apex acuminate; costa double and short or indistinct; alar cells differentiated, subquadrate to rectangular; laminal cells smooth; . Branch leaves narrowly ovate- to oblong-lanceolate, smaller to occasionally somewhat larger than stem leaves. Specialized asexual reproduction absent. Sexual condition autoicous; inner perichaetial leaves erect, oblong-lanceolate, apex subulate. Seta yellowish or reddish. Capsule erect to suberect, oblong-cylindric to oblong-ovoid, straight, not contracted below mouth; annulus differentiated or not; operculum conic to rostrate, obliquely short- to long-acute or obtuse; peristome perfect to reduced, hygroscopic; exostome teeth with external surface smooth basally, sometimes roughly papillose distally; endostome basal membrane present, segments sometimes 2-fid or fragmentary, keeled, cilia rudimentary. Calyptra naked. Spores spheric [somewhat tetrapozoidal by endosporic protonemata], finely papillose.

Species 16 (4 in the flora): North America, Mexico, Central America, South America (Andes Mountains), Europe, Asia.

Pylaisia has yellowish brown stems 2 cm long and reddish capsules; the exothecial cell walls are thin. The setae are sinistrorse-twisted basally and dextrorse-twisted distally when dry; the capsules have few phaneropore stomata on the base of the neck; and the exostome teeth are lance-subulate to blunt, distinctly lamellate, and internally trabeculate.

SELECTED REFERENCE Arikawa, T. 2004. A taxonomic study of the genus Pylaisia (Hypnaceae, Musci). J. Hattori Bot. Lab. 95: 71-154.

1 Endostome free, segments neither 2-fid nor perforate; spores usually smaller than 16 µm; stem leaves usually longer than 1.3 mm   (2)
+ Endostome wholly or partially adherent to exostome teeth, segments split, 2-fid; spores usually larger than 16 µm; stem leaves usually shorter than 1.2 mm   (3)
2 (1) Branches subjulaceous; stem and branch leaves slightly differentiated; alar cells 8-10 along margins; endostome 0.3 mm beyond mouth.   1 Pylaisia polyantha
+ Branches ± complanate-foliate, partly subjulaceous; stem and branch leaves well differentiated; alar cells 10-15 along margins; endostome 0.4-0.5 mm beyond mouth.   2 Pylaisia steerei
3 (1) Endostome free distally; stems regularly pinnate; alar cells 20-25 along margins.   3 Pylaisia selwynii
+ Endostome wholly adherent; stems irregularly pinnate; alar cells 6-12 along margins.   4 Pylaisia intricata

Lower Taxa


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