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FNA | Family List | FNA Vol. 2 | Ephedraceae | Ephedra

2. Ephedra aspera Engelmann ex S. Watson, Proc. Amer. Acad. Arts. 18: 157. 1883.

Boundary ephedra, pitamoreal

Ephedra nevadensis S. Watson var. aspera (Engelmann ex S. Watson) L. D. Benson; E. peninsularis I. M. Johnston; E. reedii Cory

Shrubs erect, 0.5--1.5 m. Bark gray, cracked and fissured. Branches opposite or whorled, rigid, angle of divergence about 30°. Twigs pale to dark green, becoming yellow with age, not viscid, slightly to strongly scabrous, with numerous longitudinal grooves; internodes 1--6 cm. Terminal buds conic, 1--2 mm, apex obtuse. Leaves opposite (rarely in whorls of 3), 1--3(--5) mm, connate to 1/2--7/8 their length; bases thickened, brown, shredding with age, ± persistent; apex obtuse. Pollen cones 2 (rarely 1 or whorled) at node, obovoid, 4--7 mm, sessile or rarely on short peduncles; bracts opposite, 6--10 pairs, yellow to red-brown, obovate, 3--4 × 2--3 mm, membranous; bracteoles slightly exceeding bracts; sporangiophores 4--5 mm, 1/2 exserted, with 4--6 sessile to short-stalked (less than 1 mm) microsporangia. Seed cones usually 2 at node, ovoid, 6--10 mm, sessile or on short, scaly peduncles; bracts opposite, 5--7 pairs, circular, 4--7 × 2--4 mm, membranous, with red-brown thickened center and base, margins entire. Seed 1, ellipsoid, 5--8 × 2--4 mm, light brown to brown, smooth to slightly scabrous.

Coning March--April. Dry rocky slopes, ravines, and fans; 500--1800 m; Ariz., Calif., N.Mex., Tex.; n Mexico.


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