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FNA | Family List | FNA Vol. 2 | Dryopteridaceae | Polystichum

7. Polystichum imbricans (D. C. Eaton) D. H. Wagner, Pteridologia. 1: 50. 1979.

Imbricate sword fern

Aspidium munitum Kaulfuss var. imbricans D. C. Eaton, Ferns N. Amer. 1: 188, plate 25, fig. 3. 1878; Polystichum munitum (Kaulfuss) C. Presl subsp. imbricans (D. C. Eaton) Munz; P. munitum var. imbricans (D. C. Eaton) Maxon

Stems ascending to erect. Leaves erect to arching back at tip, 2--8 dm; bulblets absent. Petiole 1/4--1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. Blade linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. Pinnae oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2--4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. Indusia entire to sharply dentate. Spores dark brown.

Subspecies 2: only in the flora.

Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P . californicum and P . scopulinum (D. H. Wagner 1979). Relationships to P . munitum are discussed under that species.

1 Pinnae oblong, less than 5 times longer than wide, adaxial surfaces facing upward and twisted out of plane of blade; leaves stiffly erect to tip; at base of boulders or in cliff crevices in exposed sites; California to British Columbia.   7a subsp. imbricans
+ Pinnae narrowly lanceolate, more than 5 times longer than wide, generally in 1 plane; leaf tips arching back; on forest floor in shade; only in California.   7b subsp. curtum

Lower Taxa


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