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FNA | Family List | FNA Vol. 26 | Liliaceae | Allium

64. Allium amplectens Torrey, Pacif. Railr. Rep. 4(5): 148. 1857.

Allium acuminatum Hooker var. gracile Alph. Wood; A. attenuifolium Kellogg; A. attenuifolium var. monospermum (Jepson) Jepson; A. monospermum Jepson; A. occidentale A. Gray; A. serratum S. Watson

Bulbs 1–15+, increase bulbs absent or ± equaling parent bulbs, never appearing as basal cluster, not clustered on stout primary rhizome, ovoid to ± globose, 0.6–1.5 × 0.6–1.3 cm; outer coats enclosing 1 or more bulbs, brown, prominently cellular-reticulate, membranous, cells in ± vertical rows, forming irregular herringbone pattern, transversely elongate, V-shaped, without fibers; inner coats usually dark red, sometimes white to pink, cells obscure, quadrate. Leaves persistent, withering from tip at anthesis, 2–4, basally sheathing, sheaths not extending much above soil surface; blade solid, subterete or ± channeled, 10–36 cm × 0.5–2 mm, margins entire. Scape persistent, solitary, erect, solid, terete, 15–50 cm × 3–5 mm. Umbel shattering after seeds mature, each flower deciduous with its pedicel as a unit, erect, compact, 10–50-flowered, hemispheric, bulbels unknown; spathe bracts persistent, 2–3, 6–13-veined, ovate, ± equal, apex short-acuminate. Flowers stellate, 5–9 mm; tepals spreading at anthesis, white to pink, lanceolate, ± equal, becoming papery and connivent over capsule, margins entire, apex acute; stamens included; anthers yellow or purple; pollen yellow; ovary crested; processes 6, lateral, ± prominent, ± rectangular, margins entire; style linear, equaling stamens; stigma capitate, scarcely thickened, unlobed; pedicel 4–16 mm. Seed coat dull; cells minutely roughened. 2n = 14, 21, 28.

Flowering Apr--Jul. Clay soils, including serpentine, dry slopes, and open plains; 0--1800 m; B.C.; Calif., Oreg., Wash.

All three chromosome races of Allium amplectens are widespread. The triploids are achiasmatic, causing a breakdown in the first meiotic division. This is followed by a normal second division resulting in pollen dyads that are, presumably, nonfunctional; seeds are produced by apomixis. The diploids and tetraploids produce normal pollen, in tetrads, that appears to be functional.


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