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FNA | Family List | FNA Vol. 23 | Cyperaceae | Isolepis

3. Isolepis cernua (Vahl) Roemer & Schultes in J. J. Roemer et al., Syst. Veg. 2: 106. 1817.

Scirpus cernuus Vahl, Enum. Pl. 2: 245. 1805; S. cernuus var. californicus (Torrey) Beetle; S. cernuus subsp. californicus (Torrey) Thorne

Plants annual (or perennial?); rhizomes usually obscured by culm bases and very short, sometimes vertical and elongated. Culms 4–40 cm × 0.2–0.5 mm. Leaves sometimes sparsely orange-punctate at 10–15X; sheaths usually reddish proximally; distal blade rudimentary to much longer than sheath, often exceeding culm, to 20 cm × 0.2–1 mm. Inflorescences: involucral bract 1, sometimes subtending flower or resembling enlarged floral scale, 2–6(–23) mm. Spikelets 2–5 × 1–2 mm; scales partly or completely dark orange to red-brown, rarely stramineous, midrib greenish to stramineous, not gibbous, obscurely to prominently 3–11-veined, midrib keeled near apex, membranous, hyaline, apex rounded to acute, with mucro less than 0.1 mm; proximal scale to 2 mm; other scales 1.2–1.8 × 1–1.3 mm. Flowers: anthers 0.3–0.6 mm; styles 3-fid or 3-fid and 2-fid. Achenes falling separately from scales, medium to dark brown or stramineous, ellipsoid to obovoid, compressed-trigonous to thickly biconvex, lateral angles usually prominent, abaxial angle prominent to obscure, faces convex or adaxial face slightly concave, 0.8–1 × 0.5–0.7 mm, distinctly papillose at 10–15X to obscurely papillose at 40X, often with thin whitish surface layer. 2n = 30.

Fruiting late spring–winter (Pacific Coast), winter–spring (Texas). Wet, freshwater to brackish places on beaches, dunes, marine bluffs, sandy areas, mostly coastal; 0–800 m; B.C.; Calif., Oreg., Tex., Wash.; Mexico (Baja California); temperate South America; Eurasia; Africa; Australia; New Zealand.

Isolepis cernua is widespread and variable. Four varieties were recognized by A. M. Muasya and D. M. Simpson (2002). Only var. ceruna is known from North America. The earliest collection I have seen from the Pacific Coast is from 1888; the earliest collection I have seen from Texas is from 1974.


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