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FNA | Family List | FNA Vol. 4 | Cactaceae | Grusonia

6. Grusonia clavata (Engelmann) H. Robinson, Phytologia. 26: 176. 1973.

Club-cholla

Opuntia clavata Engelmann in F. A. Wislizenus, Mem. Tour N. Mexico, 95. 1848; Cactus clavatus (Engelmann) Lemaire; Corynopuntia clavata (Engelmann) F. M. Knuth

Shrubs, forming mats, widely spreading, 5-15 cm, in series of usually 1-2 stem segments. Roots diffuse. Stem segments short-clavate, strongly narrowed at base, 2.5-5(-7.5) × 1.5-3 cm; tubercles prominent, oval, 5-10(-16) mm, narrow, 4-6 times longer than wide, obscured by interlacing spines; areoles 3 mm in diam.; wool white to gray. Spines 7-15 per areole, primarily in distal areoles; major 3-5 abaxial spines deflexed, white, flattened, longest central spine daggerlike, broadly tapered, 12-35 × 1.5+ mm basally; major 1-3 adaxial spines ascending, white, yellow to tan, angular-flattened to subterete. Glochids in adaxial 1/4 of areole, yellowish white, ± 4 mm. Flowers: inner tepals bright yellow, to 25 mm; filaments light yellow-green to white; style and stigma lobes white. Fruits yellow, barrel-shaped to ellipsoid, 30-45 × 15-25 mm, fleshy, spineless, densely ± yellow-glochidiate; areoles 35-55. Seeds yellowish white, 4.5 × 4 mm, smooth. 2n = 22.

Flowering late spring (May-Jun). Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils; 1800-2500 m; N.Mex.

L. D. Benson (1982) mapped "Opuntia clavata" in northeast Arizona, but no supporting specimens have been found. The report may be based on low-growing specimens of Cylindropuntia whipplei, a cholla common in that area.


 

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