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FNA | Family List | FNA Vol. 28 | Orthotrichaceae | Ulota

9. Ulota phyllantha Bridel, Muscol. Recent., suppl. 4: 113. 1818.

Orthotrichum fasciculare Bridel; Ulota maritima Müller Hal. & Kindberg

Plants 0.6-5.5 cm. Stems erect. Stem leaves contorted-crisped when dry, oblong, lanceolate, or linear-lanceolate, 2-3.5 mm; base ovate; margins revolute proximally, narrowly recurved distally; apex acute; basal laminal cells linear-elongate; distal cells 7-10 µm, papillae conic, large. Specialized asexual reproduction by gemmae at leaf apices. Sexual condition dioicous; perichaetial leaves somewhat longer than stem leaves. Seta 4-5 mm. Capsule oblong-cylindric, 2-2.5 mm, 8-ribbed to 1/2 length, mouth small, not smaller than mid capsule; stomata unknown; mature peristome unknown; exostome teeth erect, reflexed with age, papillose; endostome unknown. Calyptra conic, moderately hairy. Spores unknown.

Tree branches and trunks along coast, associated with salt-spray; low elevations; B.C., N.B., Nfld. and Labr., N.S., Que.; Alaska, Calif., Maine, Oreg., Wash.; s South America; Europe; Africa.

Ulota phyllantha twisted-contorted leaves that end in a short-excurrent costa usually covered by dark brown elongate gemmae. This species is dioicous and rarely has capsules, unlike other species of the genus that are almost always with sporophytes. No other species of Ulota has abundant gemmae; however, even when these are dispersed the stoutly excurrent costa forming a short cusp and the maritime habitat distinguish the species. Ulota phyllantha is a species of both coasts, as it is known from northern California north to southern Alaska and the Alaskan Peninsula in the West and from Maine north to southern Labrador in the East.


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