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FNA | Family List | FNA Vol. 8 | Saxifragaceae | Lithophragma

8. Lithophragma heterophyllum (Hooker & Arnott) Hooker & Arnott in J. Torrey and A. Gray, Fl. N. Amer. 1: 584. 1840.

Hillside woodland star

Tellima heterophylla Hooker & Arnott, Bot. Beechey Voy., 346. 1839; Lithophragma trilobum Rydberg

Plants slender. Flowering stems simple, 20-40 cm. Leaves in basal rosette and cauline, basal shallowly 2-5-lobed, cauline (2-10), 3-lobed with secondary teeth, usually much reduced, similar to basal, (bearing axillary bulbils, aerial bulbils may replace flowers in axils); stipules small, not decurrent onto petiole, (margins fimbriate); petiole to 8 cm; blade dark green or reddish green, orbiculate, (margins toothed), surfaces hairy. Inflorescences 1-4(-6), (lax), nodding, 2-6(-12)-flowered racemes, often many-branched, (15-50 cm). Pedicels equaling or shorter than hypanthium, (flowers long-pedicellate). Flowers persistent, not fragrant, horizontal; hypanthium narrowly campanulate with truncate base, not elongating in fruit, throat open, (length 2 times diam.); sepals erect in bud, becoming widely spreading after anthesis, triangular; petals (completely exserted), widely spreading, (lax), white, obovate-pandurate, lamina spoon-shaped, narrowly clawed, unlobed or 3(-5)-lobed, 5-12 mm, ultimate margins entire; ovary superior; styles much exserted in fruit; stigma papillae apical. Seeds 0.4-0.5 mm, tuberculate (tubercles in 3-19 rows, blunt). 2n = 14.

Flowering Jun. Partly or fully shaded, well-drained slopes of oak or mixed coniferous-oak woodland; 0-1500 m; Calif., Oreg.

Lithophragma heterophyllum is known in the coastal mountains of southern Oregon and in California from Humboldt to Los Angeles counties. Its variable morphology is associated with the amount of available moisture and site exposure.

Sympatric populations of Lithophragma heterophyllum and L. bolanderi occur in the south-central California Coast Ranges. The distinguishing characteristics for L. heterophyllum are hypanthium shape, petal outline, and exsertion of stigma and style in fruit.


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