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40. Cirsium occidentale (Nuttall) Jepson, Fl. W. Calif. 509. 1901.
Western thistle
Carduus occidentalis Nuttall, Trans. Amer. Philos. Soc., n. s. 7: 418. 1841
Biennials, 5–400 cm; taproots. Stems usually 1, thinly to densely gray- or white-tomentose, sometimes ± glabrate; branches few–many, usually from above mid or near base in compact, moundlike dwarf plants, ascending to spreading. Leaves: blades oblong–elliptic to oblanceolate, 6–40 × 1.5–10+ cm, shallowly to deeply pinnatifid, lobes usually rigidly spreading, undivided or with 1–2 pairs of coarse teeth or lobes, main spines 5–15 mm, both faces gray- to white-tomentose, sometimes ± glabrate or adaxial faces green, thinly arachnoid-tomentose; basal sometimes present at flowering, petiolate or sessile and bases tapered. spiny-winged; principal cauline much reduced distally, sessile, bases decurrent or not, as spiny wings; distal much reduced, linear, ± bractlike. Heads 1–many in loose to tight clusters (barely raised above rosette in dwarf plants). Peduncles 1–30 cm. Involucres ovoid to spheric, 1.5–5 × 1.5–8 cm, arachnoid to ± loosely tomentose, often adjacent phyllaries connected by conspicuous arachnoid trichomes, sometimes glabrous or glabrate. Phyllaries in 7–10 series, subequal to strongly imbricate, green or stramineous to purple-tinged, linear to narrowly lanceolate, abaxial faces without glutinous ridge; outer and mid bodies appressed, entire, apices deflexed to spreading or ascending, short-triangular to elongate, linear-acicular, spines spreading to reflexed, 1–10+ mm; apices of inner erect, often flexuous, flat. Corollas white to lavender, pink, rose-purple, or red, 18–40 mm, tubes 8–18 mm, throats 5–7 mm, lobes 5–10 mm; style tips 4–5 mm. Cypselae ± brown, 5–6 mm, apical collars not differentiated; pappi 15–30 mm.
Varieties 7 (7 in the flora): w United States.
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1 |
Plants compact, rounded, moundlike; heads usually not much elevated above leaves |
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40c Cirsium occidentale var. compactum |
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Plants usually erect; principal heads usually conspicuously pedunculate |
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(2) |
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2 (1) |
Corollas white to light purple or rose |
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40d Cirsium occidentale var. californicum |
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Corollas deep purple to bright pink or red |
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(3) |
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3 (2) |
Plants densely white-tomentose; phyllaries persistently white-tomentose (except spines); outer phyllaries usually very long, spreading to reflexed |
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40g Cirsium occidentale var. candidissimum |
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Plants variably tomentose, sometimes ± glabrate; phyllaries ± arachnoid to floccose-tomentose, sometimes green and glabrate; outer phyllaries short to long, ascending to spreading or reflexed |
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(4) |
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4 (3) |
Involucres usually about as long as wide or wider than long; phyllaries densely and persistently arachnoid with fine trichomes connecting tips of adjacent phyllaries |
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(5) |
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Involucres usually longer than wide; phyllaries tomentose or glabrate, sparingly or not arachnoid with fine trichomes connecting tips of adjacent phyllaries |
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(6) |
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5 (4) |
Phyllary apices ± imbricate, the proximal usually shorter than medial and distal, lanceolate to linear-acicular, 0.5–15 mm; co- rollas bright purple |
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40a Cirsium occidentale var. occidentale |
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Phyllary apices subequal, all long- acicular, 1.5–3 cm; corollas light to deep reddish purple |
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40b Cirsium occidentale var. coulteri |
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6 (4) |
Corollas 20–24 mm, deep reddish purple; s Santa Lucia Mountains of San Luis Obispo County, California |
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40e Cirsium occidentale var. lucianum |
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Corollas 23–35 mm, bright pink to red; widespread |
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40f Cirsium occidentale var. venustum |
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