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FNA | Family List | FNA Vol. 21 | Asteraceae | Harmonia

1. Harmonia nutans (Greene) B. G. Baldwin, Novon. 9: 464. 1999.

Callichroa nutans Greene, Pittonia 2: 227. 1892; Layia nutans (Greene) Jepson; Madia nutans (Greene) D. D. Keck

Plants 5–25 cm; proximal un-branched portions of primary stems shorter or longer than branches supporting heads. Leaves ± evenly distributed on stems. Heads usually reflexed in bud and fruit. Phyllaries 4–8, hirsute and/or hirtellous near folded edges. Ray florets 4–8; laminae 3–7 mm. Disc florets 7–30, bisexual, fertile. Ray cypselae not gibbous, beakless; pappi 0. Disc cypselae 2.3–4.5 mm; pappi of 9–11 lance-attenuate, fimbrillate scales 2–3.7 mm. 2n = 18.

Flowering Apr–Jun. Rocky soils, openings in chaparral or woodlands; of conservation concern; 100–1000 m; Calif.

Harmonia nutans occurs in the southern North Coast Ranges and northern San Francisco Bay area, mostly on volcanic substrates. Based on molecular phylogenetic data, H. nutans evidently represents the sister-group to the serpentine-endemic species of Harmonia (B. G. Baldwin 2001).


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