Crystal Springs lessingia

Lessingia hololeuca Greene var. arachnoidea (Greene) J. T. Howell; L. micradenia Greene var. arachnoidea (Greene) Ferris

Plants 15–80 cm. Stems erect, tan, glabrous or villous. Leaves: basal withering by flowering; cauline margins entire, faces eglandular, abaxial glabrous or villous. Heads borne singly, at ends of branchlets. Involucres obconic, 4–8 mm. Phyllaries purple-tipped, faces arachnose, sometimes gland-dotted; inner scarious. Disc florets (3–)8–18; corollas pink to lavender (color more intense in tubes); style-branch appendages truncate-penicillate, 0.1–0.3 mm. Pappi tan, usually shorter than cypselae, sometimes forming coronas, rarely equaling or longer than cypselae (Sonoma County). 2n = 10.

Flowering Jul–Oct. Grasslands, coastal scrub, chaparral, woodlands, serpentinite soils; of conservation concern; 100–300 m; Calif.

Lessingia arachnoidea is known from near Crystal Springs Reservoir in San Mateo County and near Camp Meeker in Sonoma County. Previous circumscriptions of L. arachnoidea (pappi shorter than cypselae, forming coronas) have been expanded to include plants (from Sonoma County) with pappi equaling or longer than cypselae. Alternatively, the plants with longer pappi could have been accommodated in L. ramulosa (as a glandless form); the absence of basal leaves at flowering and the presence of arachnoid indument on the phyllaries suggest a closer affinity to L. arachnoidea.