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FNA | Family List | FNA Vol. 19, 20 and 21 | Asteraceae | Psilocarphus

5. Psilocarphus chilensis A. Gray in A. Gray et al., Syn. Fl. N. Amer. ed. 2. 1: 448. 1886.

Round woolly marbles

Micropus globiferus Bertero ex de Candolle in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 460. 1836, not Psilocarphus globiferus Nuttall 1840; P. tenellus Nuttall var. globiferus (Bertero ex de Candolle) Morefield; P. tenellus var. tenuis (Eastwood) Cronquist

Plants mostly greenish, thinly arachnoid-sericeous (in coastal forms grayish to whitish, ± lanuginose). Stems mostly (1–)2–7, ascending to ± prostrate; proximal internode lengths (2–)3–6 times leaf lengths. Capitular leaves erect to incurved, appressed to heads, ovate to broadly elliptic, widest in proximal 2/3, longest 5–12 mm, lengths mostly 1.2–1.8(–2) times widths, 1–2(–2.5) times head heights. Heads ± spheric, largest 3–5.5 mm. Receptacles unlobed. Pistillate paleae usually individually visible through indument, longest mostly 1.5–2.7 mm. Staminate corollas 0.8–1.3 mm, lobes mostly 4. Cypselae narrowly obovoid, somewhat compressed, 0.6–1.2 mm.

Flowering and fruiting mid Mar–early Jul. Saturated to drying vernal pool margins, seasonally inundated sites, coastal interdune areas; 0–600 m; Calif.; South America (Chile).

Psilocarphus chilensis occurs mainly in west-central California and central Chile; one recent collection is from southern California (western Riverside County). Ecotypes from coastal interdune areas are more lanuginose with shorter stems and internodes than intergrading populations farther inland; they are indistinguishable from the type of Micropus globiferus from Chile (J. D. Morefield 1992d). Psilocarphus chilensis and P. tenellus are at least as distinct as the other species of Psilocarphus; contrary to suggestions by A. Cronquist (1950), intermediates between the two are at most very uncommon.

Psilocarphus berteri I. M. Johnston is a superfluous name for P. chilensis. I. M. Johnston (1938) erroneously applied P. chilensis to a species not including the type of Micropus globiferus; such plants are here included in P. brevissimus var. brevissimus.


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