6b. Chaenactis douglasii (Hooker) Hooker & Arnott var. alpina A. Gray in A. Gray et al., Syn. Fl. N. Amer. 1(2): 341. 1884.
Alpine dustymaidens or pincushion
Chaenactis alpina (A. Gray) M. E. Jones; C. alpina var. leucopsis (Greene) Stockwell; C. alpina var. rubella (Greene) Stockwell; C. panamintensis Stockwell
Perennials, mostly (2–)5–10(–20) cm (cespitose to ± matted). Stems (1–)10–25+. Leaves strictly basal, to (1–)2–6 cm, indument persistent or glabrate. Heads 1(–2) per stem. Phyllaries: longest 9–12 mm; outer ± stipitate-glandular (rarely eglandular) and, often, arachnoid to lanuginose. 2n = 12.
Flowering Jul–Sep. Rocky or gravelly alpine ridges, talus, fell-fields, rock crevices; 2700–4000 m; Calif., Colo., Idaho, Mont., Oreg., Utah, Wyo.
In California, relatively few populations of Chaenactis douglasii approach typical var. alpina. A report of var. alpina from the Spring Mountains of Clark County, Nevada (P. Stockwell 1940, as C. panamintensis), could not be confirmed and was likely based on stunted var. douglasii; it should be sought in the Carson, Ruby, and Snake ranges of northern Nevada. A report of var. alpina from southern British Columbia has not been verified by me. Variety alpina is sometimes cultivated, especially for rock gardens, and it may be encountered outside its native range.
Local populations of these largely alpine plants can appear highly distinctive and uniform, leading to their past recognition as a separate species. Across the range, most such populations intergrade continuously with local populations of var. douglasii, as is often evident among specimens of the same gathering. The relative development of roots, stems, and branches appears highly mutable throughout C. douglasii.
Chaenactis pumila Greene was assigned to the variety or species alpina by R. D. Dorn (1988c), P. Stockwell (1940), and others; it is a small form of var. douglasii, as assigned by D. C. Eaton in S. Watson (1871).