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FNA | Family List | FNA Vol. 20 | Asteraceae | Heterotheca

8a. Heterotheca fulcrata (Greene) Shinners var. fulcrata

Chrysopsis cryptocephala Wooton & Standley; C. elata Osterhout; C. resinolens A. Nelson; C. resinolens var. ciliata A. Nelson; C. viscida subsp. ciliata (A. Nelson) S. F. Blake

Stems sparsely to moderately hispido-strigose and stipitate-glandular. Distal cauline leaf blades ovate to lanceolate, (12–)16–39(–55) × 5–15 mm, margins with few long-hispido-strigose cilia, faces sparsely to moderately hispido-strigose and stipitate-glandular. Subtending bracts ovate, usually large (7–22 × 2–6 mm), leaflike, well differentiated. Involucres 6.7–10.3 mm. Phyllaries lanceolate to triangular-lanceolate, moderately strigose and stipitate-glandular. Ray laminae (8–)9.8–14.5 mm. Cypsela faces sparsely to moderately strigose. 2n = 18, 36.

Flowering (Jun–)Aug–Sep(–Oct). Riparian zones in ponderosa pine and Douglas fir forests, grassy clearings in Ponderosa pine forest, lower spruce-fir zone, sandy riverbanks, roadsides, coarse sandy granitic soils with gravel, crushed lava gravelly soils, moist disturbed soils, shale slopes with sandstone outcrops, open, upper grass and sagebrush slopes; 1500–3100 m; Ariz., Colo., Idaho, Nev., N.Mex., Tex., Utah, Wyo.

Variety fulcrata grows in the Front Range of the Rocky Mountains (Colorado and Wyoming), and in disjunct populations in Arizona, New Mexico, and Utah; it is considered very rare and atypical in adjacent Idaho, Nevada, and Texas, where it is of conservation concern. Variety fulcrata is the most variable member of Heterotheca fulcrata. It is characterized by having more hairs than glands per area of leaf face, the hairs being sparse to moderate. Many collections are morphologically intermediate in indument features with var. amplifolia. A few intermediates with the other two varieties also have been collected, as well as a number of possible hybrids with H. pumila and with several varieties of H. villosa.


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