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FNA | Family List | FNA Vol. 20 | Asteraceae | Heterotheca

15f. Heterotheca villosa (Pursh) Shinners var. minor (Hooker) Semple, Novon. 4: 54. 1994.

Chrysopsis villosa (Pursh) Nuttall ex de Candolle var. minor Hooker, London J. Bot. 6: 244. 1847; C. arida A. Nelson; C. asprella Greene; C. bakeri Greene; C. columbiana Greene; C. compacta Greene; C. floribunda Greene; C. grandis Rydberg; C. hirsuta Greene; C. hirsutissima Greene; C. hispida (Hooker) de Candolle; C. villosa (Pursh) Nuttall ex de Candolle var. discoidea A. Gray; C. villosa var. hispida (Hooker) A. Gray; C. wisconsinensis Shinners; Diplogon villosum (Pursh) Kuntze var. discoideum (A. Gray) Kuntze; Diplopappus hispidus Hooker; Heterotheca villosa (Pursh) Shinners var. hispida (Hooker) V. L. Harms

Stems decumbent to ascending-erect, (9–)16–33(–48) mm, sparsely to densely hispido-strigose, sparsely to abundantly long-hirsute, sparsely to densely stipitate-glandular. Distal cauline leaf blades usually narrowly to broadly oblanceolate or oblong, sometimes ovate (in mountains), (11–)16–28(–40) mm, ± reduced distally, bases usually narrowly to broadly convex-cuneate to attenuate, sometimes rounded, margins flat, sometimes remotely undulate, apices obtuse or acute, faces sparsely to moderately hispido-strigose, (1–75/mm 2 ), sparsely to moderately glandular (glands 1–32/mm2 ). Heads 1–13(–42) in congested to open, corymbiform arrays, branches usually not very long. Peduncles 4.5–40(–60) mm, moderately to densely hispido-strigose, usually sparsely to moderately stipitate-glandular rarely eglandular; bracts usually linear-oblong, reduced, rarely linear-lanceolate and leaflike, sparsely to moderately hispido-strigose, sparsely to moderately stipitate-glandular. Involucres cylindric to campanulate (fresh), (5.5–)6.5–8.5(–9.3) mm. Phyllaries narrowly triangular-lanceolate, faces usually sparsely, rarely densely strigose (hairs to 1 mm), usually sparsely to moderately stipitate-glandular, rarely eglandular. Ray florets (7–)10–18(–26), laminae (4–)6–10(–12.6) mm. 2n = 18, 36.

Flowering (May–)Jun–Aug(–Nov). Gravelly, sandy, and loamy soils, crevices in granite, limestone rocks, marble rocks, eroded granites and sandstones, basaltic cliffs, lava flows, rocky slopes, dry ledges, stream banks, glacial outwashes, roadside railroad embankments, grasslands, ponderosa pine-oak woods, open pinyon-juniper associations; (60–)300–2900(–3200) m; Alta., B.C., Man., Sask.; Ariz., Calif., Colo., Idaho., Ill., Iowa, Kans., Minn., Mont., Nebr., Nev., N.Mex., N.Dak., Oreg., S.Dak., Utah, Wash., Wis., Wyo.

Variety minor is distinguished by its usually narrowly to broadly oblancelate distal cauline leaves (sometime oblong or ovate) and moderately glandular and hispido-strigose indument. The leaf bases are usually narrowly to broadly cuneate or attenuate (rarely rounded). Distal leaf faces usually have about 10–50 hairs/mm 2 and about 4–20 glands/mm 2 . The variety is the most variable in the species and includes some local distinctive morphotypes that grade into other forms. Plants intermediate between this and all other varieties occur in areas where the ranges are sympatric, and they make infraspecific taxonomy of the species difficult. The variety has been incorrectly referred to as var. hispida (a later synonym) in most floras.

The status of Heterotheca barbata (Rydberg) Semple (Chrysopsis barbata Rydberg), the Spokane goldenaster, is uncertain. J. C. Semple (1996) treated it as a separate species to draw attention to the problem; a detailed description based on the type and detailed illustrations were included. It is known from the type collection along the Spokane River Valley east of Spokane, Idaho, and two down-river, atypical collections (Benton and Spokane counties, Washington). It flowers in July (sometimes August). It is similar to H. villosa var. minor, but differs in having lanceolate-elliptic distal cauline leaves (34–38 × 8–9 mm) that are little reduced distally, long branches (each with one to a few large heads), and disc corollas with a few, very long hairs on the tube. Further work is needed to increase the number of specimens available for a detailed comparison with H. villosa var. minor in order to clarify whether H. barbata warrants species level recognition, should be included in H. villosa as a variety, or placed in synonymy under var. minor.


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