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FNA | Family List | FNA Vol. 7 | Brassicaceae | Thelypodium

5. Thelypodium howellii S. Watson, Proc. Amer. Acad. Arts. 21: 445. 1886.

Cartiera howellii (S. Watson) Greene; Thelypodiopsis howellii (S. Watson) O. E. Schulz

Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles). Stems branched distally, 1-8(-9.3) dm. Basal leaves: petiole 0.5-2 cm, ciliate; blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2-10(-13.5) cm × 10-23(-40) mm, margins often lyrate to dentate, sometimes entire or repand. Cauline leaves (ascending); sessile; blade often lanceolate to linear, sometimes oblong, (0.8-)1-9.7(-14.5) cm × 2-10(-30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire. Racemes lax, elongated in fruit, (flower buds ovate to lanceolate). Fruiting pedicels usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5-)3.5-8(-14.5) mm, not flattened at base. Flowers: sepals erect, linear or lanceolate, (3.5-)5-8.5(-11.5) × (0.7-)1-2(-2.5) mm; petals lavender to purple, linear or oblanceolate to spatulate, (6-)9-16.5(-22) × 0.5-2.7(-3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5-)4-7(-11) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments slightly tetradynamous, median pairs (3-)5.5-9(-12) mm, lateral pair (2.5-)4.5-8(-10) mm; anthers slightly exserted to included, linear, (1.5-)2-4(-5.5) mm, not circinately coiled; gynophore (stout), 0.5-1(-3.5) mm. Fruits often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5-6.5(-8.2) cm × 1-1.3(-2) mm; ovules 22-40 per ovary; style cylindrical or subclavate, (0.5-)1-2.7(-4.3) mm. Seeds (1-)1.2-1.7 (-2) × 0.5-0.7 mm.

Subspecies 2 (2 in the flora): w United States.


1 Petals usually linear to narrowly oblanceolate, rarely spatulate, 0.5-1.2(-3) mm wide; median filaments connate most of their lengths; ne California, c, s Oregon, s Washington.   5a Thelypodium howellii subsp. howellii
+ Petals usually spatulate, rarely oblanceolate, (1.2-) 1.5-2.7(-3) mm wide; median filaments distinct; e Oregon.   5b Thelypodium howellii subsp. spectabilis

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