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FNA | Family List | FNA Vol. 7 | Brassicaceae | Physaria

22. Physaria didymocarpa (Hooker) A. Gray, Gen. Amer. Bor. 1: 162. 1848.

Common twinpod

Vesicaria didymocarpa Hooker, Fl. Bor.-Amer. 1: 49, plate 16. 1830; Coulterina didymocarpa (Hooker) Kuntze

Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). Stems several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. Basal leaves (forming a strong rosette; long-petiolate); blade obovate, 1.5-4(-8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). Cauline leaves: blade oblanceolate, 1-2 cm (width 4-8 mm), margins entire or with occasional tooth, (apex acute). Racemes congested, (elongated in fruit, greatly exceeding leaves). Fruiting pedicels (spreading, straight or slightly curved), 8-12 mm. Flowers: sepals lanceolate to oblong, 6-8 mm, (often keeled); petals spatulate, 10-12 mm. Fruits (erect), didymous, inflated, 10-20 × 10-20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy; replum obovate to broadly oblong, not constricted, 3-4 mm, as wide as or wider than fruit, apex obtuse; ovules (4-)8 per ovary; style 7-9 mm. Seeds flattened.

Subspecies 3 (3 in the flora): w North America.

1 Trichomes spreading throughout; basal leaf blade margins dentate.   22b Physaria didymocarpa subsp. lanata
+ Trichomes appressed (except fruits in subsp. didymocarpa); basal leaf blade margins repand to dentate, ± lyrate or, rarely, entire   (2)
2 (1) Fruits moderately inflated, trichomes spreading.   22a Physaria didymocarpa subsp. didymocarpa
+ Fruits highly inflated (papery), trichomes appressed.   22c Physaria didymocarpa subsp. lyrata

Lower Taxa


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