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FNA | Family List | FNA Vol. 28 | Brachytheciaceae | Brachytheciastrum

6b. Brachytheciastrum velutinum var. salicinum (Schimper) Ochyra & Zarnowiec, Biodivers. Poland. 3: 172. 2003.

Brachythecium salicinum Schimper in P. Bruch and W. P. Schimper, Bryol. Europ. 6: 19, plate 548. 1853; B. suberythrorrhizon Renault & Cardot; B. velutinum var. venustum (De Notaris) Archangelico

Plants green to golden brown.

Rock and soil near rock outcrops, open to moderately shaded places; low to high elevations (0-3000 m); Ariz., Calif., Colo., Idaho, Mont., N.Mex., Oreg., Tex., Utah, Wash., Wyo.; Europe; w Asia; n Africa; Atlantic Islands.

Stems creeping to ascending and erect in dense growth, irregularly pinnate, branches terete- or rarely subcomplanate-foliate. Stem leaves erect, densely arranged, straight, often flexuose distally, lanceolate, 1.2-1.8 × 0.3-0.5 mm; margins serrate, serrulate, or subentire; laminal cells (60-)80-140 × 4-5µm. Branch leaves straight. Seta slightly rough proximally and smooth near capsule, or smooth throughout.

The molecular phylogenetic data of A. Vanderpoorten et al. (2005) and Vanderpoorten and B. Goffinet (2006) support the species status of the Brachytheciastrum velutinum group with smooth setae. However, its acceptance at the species level makes it difficult to almost impossible to identify the majority of sterile collections. Thus, the approach of most American authors is retained here (see E. Lawton 1971; D. H. Norris and J. R. Shevock 2004) until the differences in gametophytes are found between var. salicinum and var. velutinum. Although a number of characters differentiate these two varieties, in practice their identification is not easy, and this complex obviously needs further study.


 

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