19. Brachythecium udum I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim). 1908(3): 4, plate 1, fig. 1. 1908.
Brachythecium mildeanum var. udum (I. Hagen) Mönkemeyer; B. salebrosum subsp. udum (I. Hagen) J. J. Amann
Plants medium-sized to large, in dense mats, green to pale stramineous. Stems to 8 cm, ascending to upright, terete-foliate, quite loosely irregularly pinnate, often almost unbranched, branches to 7 mm, straight, terete-foliate. Stem leaves erect, occasionally erectopatent, closely imbricate, ovate to ovate-lanceolate, broadest at 1/12-1/9 leaf length, concave, not to somewhat plicate, 1.9-2.4 × 1-1.3 mm; base slightly rounded, inconspicuously decurrent; margins recurved usually only near base, entire to slightly serrulate; apex gradually tapered, often abruptly acuminate; costa to 40-70% leaf length, stronger near base, often much weaker distally, terminal spine absent; alar cells short-rectangular, enlarged, 20-30 × 20 µm, walls moderately thin, region slightly differentiated or enlarged, indistinctly or rarely distinctly delimited, of to 5 × 6 cells, ; laminal cells linear, 70-115 × 7-9 µm, ; basal cells 12-18 µm wide, region in 2 or 3 rows. Branch leaves smaller, narrower. Sexual condition dioicous. Sporophytes unknown.
Wet soil, rock over soil, Arctic and alpine habitats; low elevations (100 m); Alaska; n Eurasia.
Brachythecium udum occurs in Siberia and northern Europe; it has been found in collections by W. C. Steere from Alaska (No Luck Lake), and probably occurs in northern Canada. The Alaska specimen was identified as B. albicans, which is superficially rather similar, but the different patterns of basal areolation easily distinguish them. Alar cells of B. udum are more differentiated than in B. acutum, and in permanently wet habitats, the alar cells can be rather strongly enlarged. Poor branching and commonly upright growth are also characteristic of B. udum.