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FNA | Family List | FNA Vol. 28 | Brachytheciaceae | Homalothecium

7. Homalothecium nevadense (Lesquereux) Renauld & Cardot, Bot. Gaz. 13: 202. 1888.

Hypnum nevadense Lesquereux, Mem. Calif. Acad. Sci. 1: 33. 1868

Plants medium-sized, in dense tufts, light green when young, golden or brownish yellow with age. Stems to 10 cm, densely regularly pinnate, branches 5-7 mm, curved to almost circinate. Stem leaves erect-appressed, triangular-lanceolate, (1-)1.3-2(-2.5) × (0.2-)0.5-0.7 (-0.8) mm; base short-rounded, narrowly decurrent; margins plane or often recurved at places, serrulate or subentire; apex acuminate or gradually tapered; costa to 70-95% leaf length, terminal spine present; alar cells ovate, 6-7 µm wide, , region of 5-10 × 6-10 cells, moderately distinctly delimited, ; laminal cells linear-flexuose, 30-85 × 5-6 µm; basal cells irregularly long-ovate, region in 1-3 rows, indistinctly delimited from distal cells. Branch leaves appressed when dry, spreading when moist, lanceolate, 1-1.7(-2.1) × 0.2-0.4 mm; margins plane or recurved at places, serrulate or serrate proximally and in alar region, entire or minutely serrulate distally; apex narrowly acute to acuminate; costa to 75-95% leaf length, variable within one shoot, terminal spine present; alar cell shape irregular, 5-10 µm wide, , region of 2-10 × 3-8 cells, distinctly delimited; laminal cells linear-flexuose, 25-65 4-6 µm; basal cells in 1 (or 2) rows; distal cells smooth. Sexual condition phyllodioicous or dioicous. Seta 0.7-1.5 cm, rough proximally, sometimes slightly so, smooth distally. Capsule erect, narrowly cylindric, straight to slightly curved, 2-2.5 mm; annulus separating by fragments; operculum long-conic to rostrate; peristome hygrocastique; exostome teeth with long transitional zone; endostome basal membrane short, segments as long as exostome teeth, very narrow, cilia absent or less than 1/4 segment length. Spores 9-16 µm.

Calcareous rock, quartzite, granite, rotten logs, trunks, soil, forests, open areas; low to high elevations (0-3100 m); Alta., B.C.; Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Homalothecium nevadense has stem leaf costae that are stout to the end, as does H. aeneum; the branch leaves may be straight, homomallous, or falcate-secund; and the spores are smooth.


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