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FNA | Family List | FNA Vol. 21 | Asteraceae

276. Agnorhiza (Jepson) W. A. Weber, Phytologia. 85: 19. 1999.

[Possibly Greek agnostos, unknown, and rhiza, root, alluding to the initially unknown roots; in protologue of basionym of type species, Greene stated, "Root unknown."]

William A. Weber

Balsamorhiza Nuttall sect. Agnorhiza Jepson, Man. Fl. Pl. Calif., 1077. 1925

Perennials, 5–60(–100) cm (taproots relatively massive; caudices sometimes branched). Stems erect and branched (mostly distally), or decumbent (and seldom branched distally). Leaves basal and cauline (basal usually scalelike on flowering stems, well-formed on non-flowering shoots); alternate; petiolate; blades (3-or 5-nerved or pinnately nerved), cordate-ovate, deltate, deltate-triangular, ovate, ovate-lanceolate, or suborbiculate, bases broadly cuneate, subcordate, or truncate, margins entire or dentate to crenate, faces glabrous or hairy (often gland-dotted). Heads radiate or discoid, borne singly (terminal) or in ± racemiform arrays. Involucres campanulate to ± hemispheric, 12–40 mm diam. Phyllaries persistent, (8–)12–26+ in 2–3+ series (subequal to unequal, outer usually surpassing inner). Receptacles flat to convex, paleate (paleae persistent, conduplicate, chartaceous). Ray florets 0 or (1–)5–21+, pistillate, fertile; corollas yellow. Disc florets 50–150+, bisexual, fertile; corollas yellow, tubes shorter than cylindric throats, lobes 5, lance-deltate (style branches stigmatic in 2, barely distinct lines, appendages filiform). Cypselae ± prismatic, 3–4-angled (faces glabrous or strigillose); pappi usually persistent, coroniform (± lacerate, sometimes with 1–4 teeth or scales), sometimes 0. x = 19.

Species 5 (5 in the flora): California, nw Mexico.


Weber, W. A. 1946. A taxonomic and cytological study of the genus Wyethia, family Compositae, with notes on the related genus Balsamorhiza. Amer. Midl. Naturalist 35: 400–452.

1 Plants (5–)15–30(–45+) cm; stems decumbent, seldom distally branched; cauline leaves: blades ovate, suborbiculate, elliptic, rounded-deltate, or oblong; involucres ± campanulate, 12–25(–30+) mm diam   (2)
+ Plants (20–)40–100 cm; stems erect, usually distally branched; cauline leaves: blades deltate, ovate, or ovate-lanceolate; involucres ± hemispheric, (15–)20–40 mm diam   (3)
2 (1) Leaves glabrous (shining, finely gland-dotted, glutinous); heads held among or beyond leaves; ray florets (7–)8–12, laminae (12–)20–30(–35) mm; cypselae 7–9 mm; pappi 0 or coroniform, 0.1–0.3 mm   1 Agnorhiza bolanderi
+ Leaves silky-villous to strigillose, glabrescent; heads in axils of (and overtopped by) leaves; ray florets 5–8(–9), laminae 8–18 mm; cypselae 9–10 mm; pappi coroniform, 1–1.5+ mm   2 Agnorhiza ovata
3 (1) Ray florets 0 or 2–3   3 Agnorhiza invenusta
+ Ray florets 10–23   (4)
4 (3) Leaf blades: faces tomentulose to pilosulous (and gland-dotted); cypselae 8–12 mm; pappi (0.5–)1–2(–3) mm   4 Agnorhiza elata
+ Leaf blades: faces sparsely hispid or scabrous (and finely gland-dotted, green, shin-ing, often vernicose); cypselae ca. 6 mm; pappi 0.1–1 mm   5 Agnorhiza reticulata

Lower Taxa


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