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FNA | Family List | FNA Vol. 27 | Grimmiaceae | Codriophorus

9a. Codriophorus P. Beauvois sect. Codriophorus

Racomitrium sect. Papillosa (Kindberg) Noguchi; Trichostomum sect. Stenotrichum Chevallier

Stem sparingly or freely irregularly, dichotomously or fasciculately branched, sometimes filiform and almost unbranched, absent short, lateral tuft-like branchlets. Leaves lingulate, elliptic, broadly ovate, ovate- or oblong-lanceolate to broadly or occasionally narrowly lanceolate; margins 1-2(-4)-stratose, recurved, reflexed, incurved or plane on both sides; apices narrowly to broadly rounded-obtuse or long acuminate to acute, muticous or very rarely shortly hyaline-tipped, entire or sharply to bluntly, remotely irregularly toothed, erose-dentate, sinuate to cristate in the upper 1/5-1/4; costa strong and broad, ceasing in mid leaf to subpercurrent, in transverse-section 2-4-stratose in the distal and medial parts, weakly to strongly convex, reniform, crescent-shaped to semi-terete abaxially, flat or convex on the adaxial side, in the basal part 3-6(-7)-layered, situated at the bottom of a shallow or deeper, usually wide-angled groove, lunate, reniform or rectangular and distinctly flattened, strongly convex on the abaxial side; basal marginal laminal cells not differentiated; distal laminal cells 1- to variously 2-stratose, irregularly rounded-quadrate, subquadrate, oval to rounded-hexagonal, strongly papillose to smooth, straight-walled in the distal part.

Species 9 (6 in the flora): North America, South America (Colombia), Eurasia, n Africa, Atlantic Islands (Azores, Canary Islands, Iceland, Madeira).

Members of sect. Codriophorus are recognized by their lingulate, broadly ovate to ovate- or oblong-lanceolate leaves, isodiametric and straight-walled distal laminal cells, and costa that is broad and robust, usually over 100 ┬Ám wide at the base. The costa is situated in a shallow, wide-angled groove and is mostly 2-4-stratose in the distal and medial parts, 3-6(-7)-stratose in the basal part, and anatomically strongly differentiated.


1 Laminal cells smooth or very slightly papillose on young leaves; leaf margins entire or with a few irregular teeth at the apex.   5 Codriophorus depressus
+ Laminal cells distinctly papillose; leaf margins remotely irregularly toothed, erose-dentate, sinuate to cristate in the distal 1/5-1/4, if entire then broadly rounded-obtuse   (2)
       
2 (1) Leaves lanceolate; apex irregularly bluntly erose-dentate; costa markedly convex and crescent-shaped abaxially.   6 Codriophorus ryszardii
+ Leaves lingulate, elliptical to broadly ovate or ovate-lanceolate; apex broadly rounded-obtuse to acute, irregularly dentate to entire   (3)
       
3 (2) Costa subpercurrent, entire and only shortly forked at the tip   (4)
+ Costa extending 1/2-3/4 way up the leaf, distinctly spurred   (5)
       
4 (3) Laminal cells strongly papillose, 1- or very rarely 2-stratose; leaves unbordered.   1 Codriophorus acicularis
+ Laminal cells indistinctly papillose over the cell walls, always variously 2-stratose in the distal part; leaves distinctly bordered with fleshy and broad limbidia.   4 Codriophorus norrisii
       
5 (3) Leaf apex broadly rounded, flat to subcucullate, always entire or sometimes sinuate to remotely bluntly erose-dentate.   2 Codriophorus mollis
+ Leaf apex broadly rounded-obtuse to subacute, often tubular, strongly dentate or very rarely subentire.   3 Codriophorus aduncoides

Lower Taxa


 

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