3. Heterosavia (Urban) Petra Hoffmann, Brittonia. 60: 152. 2008.
Maidenbush [Greek hetero-, other or different from, and genus Savia] Maidenbush [Greek hetero-, other or different from, and genus Savia]
Geoffrey A. Levin
Savia Willdenow sect. Heterosavia Urban, Symb. Antill. 3: 284. 1902
Shrubs or trees, dioecious, hairy, glabrescent, hairs simple; branching not phyllanthoid. Leaves persistent, alternate, simple, all well developed; stipules persistent; blade margins entire. Inflorescences unisexual, staminate glomes, pistillate flowers solitary. Pedicels: staminate rudimentary, pistillate present. Staminate flowers: sepals 5, distinct; petals 5; nectary extrastaminal, annular, crenate [entire]; stamens 5; filaments distinct or connate basally [to 1/2 length]; connectives not extending beyond anthers; pistillode 3-divided to base or nearly so. Pistillate flowers: sepals persistent, 5, distinct; petals 5; nectary annular, crenate [entire]; pistil 3(–4)-carpellate; styles (3–)4, distinct, 2-fid 1/2 length. Fruits capsules. Seeds 2 per locule, rounded-trigonous; seed coat dry, smooth; caruncle absent.
Species 4 (1 in the flora): Florida, West Indies (Greater Antilles).
Molecular phylogenetic analyses have shown that Savia as traditionally recognized is polyphyletic and should be treated as three genera, each assigned to a different tribe (H. Kathriarachchi et al. 2005; P. Hoffmann et al. 2006; Hoffmann 2008). Heterosavia, which includes the species formerly included in the Caribbean Savia sect. Heterosavia, belongs to Phyllantheae Dumortier and is sister to Flueggea.
SELECTED REFERENCE Hoffmann, P. 2008. Revision of Heterosavia, stat. nov., with notes on Gonatogyne and Savia (Phyllanthaceae). Brittonia 60: 136–166.