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FOC | Family List | FOC Vol. 5 | Amaranthaceae

9. Achyranthes Linnaeus, Sp. Pl. 1: 204. 1753.

牛膝属 niu xi shu

Herbs or subshrubs. Stem with conspicuous nodes; branches opposite. Leaves opposite, petiolate. Spikes terminal or axillary, erect. Flowers reflexed or spreading, perfect, with 3 bracts; midvein of bracteoles spiny, long, excurrent. Tepals 4 or 5, membranous, apex awn-pointed, rigid in fruit. Stamens 5, rarely 4 or 2, shorter than perianth; filaments connate into a short cup at base, pseudostaminodes fringed; anthers 2-loculed. Ovary oblong; style linear, persistent; stigma capitate. Utricles ovoid-oblong, ovoid, or subglobose, falling off with tepals and bracteoles. Seed oblong, lenticular.

About 15 species: tropical and subtropical regions; three species in China.

The two species Achyranthes aspera and A. bidentata are highly variable. Further study might show A. longifolia to be merely a variant of A. bidentata.

Achyranthes megaphylla Y. H. Li (Guihaia 9: 26. 1989, not Standley, 1917) was described from Yunnan and A. ogatae Yamamoto (Icon. Pl. Formosan., Suppl. 3: 23. 1927) was described from Taiwan. We have seen no specimens of either taxon and are therefore unable to treat them in this account. Further revisionary study is necessary.

1 Leaves lanceolate or broadly lanceolate; pseudostaminodes inconspicuously dentate at apex.   3 A. longifolia
+ Leaves obovate, elliptic, or oblong; pseudostaminodes ciliate or serrulate at apex.   (2)
2 (1) Leaves obovate, broadly obovate, or elliptic-oblong, apex obtuse with a mucro; bracteoles spiny, 2-winged at base; pseudostaminode long fimbriate and ciliate at apex.   1 A. aspera
+ Leaves elliptic or elliptic-lanceolate, apex caudate; bracteoles spiny, 2-parted at base; pseudostaminode serrulate at apex.   2 A. bidentata

Lower Taxa


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