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7. Ainsliaea Candolle, Prodr. 7: 13. 1838.

兔儿风属 tu er feng shu

Authors: Tiangang Gao, Susana Edith Freire & D. J. Nicholas Hind

Diaspananthus Miquel.

Herbs, perennial, rarely subshrubs. Leaves alternate, often in basal rosettes or densely aggregated near median part of stems; blade linear, ovate, elliptic, or orbicular, margin entire, crenate, denticulate, dentate, or lobed. Capitula many in spikes, racemes, or panicles, sometimes nodding, radiate or discoid, homogamous; involucres cylindric; phyllaries often multiseriate, imbricate; receptacle small, glabrous, epaleate. Florets few, (1-)3(-5), bisexual, all fertile, chasmogamous or cleistogamous; chasmogamous corollas open, deeply irregularly 5-lobed, one of slits much longer than others, lobes linear, reflexed and usually coiled; cleistogamous corolla (if present) closed, tubular, shorter than pappus; anther bases long tailed, apex appendages truncate to rounded, sometimes apiculate; styles shallowly divided at apex, style branches short, abaxially with papillae, apices obtuse to acute, rounded, or truncate. Achenes terete or fusiform, ribbed, glabrous or pilose; pappus of plumose bristles or sometimes absent in chasmogamous florets, usually uniseriate, more rarely biseriate. 2n = 24, 26.

About 50 species: Afghanistan, Bangladesh, Bhutan, China, India, Indonesia, Japan, Korea, Myanmar, Nepal, Pakistan, Philippines, Thailand, Vietnam; 40 species (28 endemic) in China; one additional species is incompletely known.

See the revision of Ainsliaea by Freire (Ann. Missouri Bot. Gard. 94: 79-191. 2007).

Incompletely known species

Ainsliaea mollis Diels, Repert. Spec. Nov. Regni Veg. Beih. 12: 514. 1922.

Ainsliaea mollis was treated by Y. C. Tseng (FRPS 79: 34. 1996) as an accepted species, occurring in Sichuan and Yunnan. It resembles A. fragrans in external morphology. No material of this species was available to the present authors. Its identity is dubious.


1 Subshrubs; stems much branched.   1 A. pertyoides
+ Perennial herbs; stems not branched   (2)
       
2 (1) Leaves aggregated near or toward median part of stem or above basal part of stem   (3)
+ Leaves in basal rosette (rarely specimens with leaves above stem base in A. apteroides)   (17)
       
3 (2) Capitula 1-flowered.   2 A. lancangensis
+ Capitula 2-4-flowered   (4)
       
4 (3) Leaves lobed   (5)
+ Leaves not lobed   (7)
       
5 (4) Leaf blade shallowly pinnately lobed.   3 A. grossedentata
+ Leaf blade palmately lobed   (6)
       
6 (5) Leaf blade 5-14 × 5.5-18 cm, shallowly 5-7-lobed.   4 A. acerifolia
+ Leaf blade (1-)2.5-3.5 × (1-)2-3 cm, deeply lobed with apical lobe elongated.   5 A. apiculata
       
7 (4) Leaf blade linear to elliptic or lanceolate (0.3-1.3 cm wide)   (8)
+ Leaf blade lanceolate or ovate (1-12 cm wide)   (9)
       
8 (7) Leaf blade linear to elliptic, 3-7 mm wide, apex acute, 1-veined.   6 A. walkeri
+ Leaf blade lanceolate, 5-13 mm wide, apex acuminate, 3-veined.   7 A. trinervis
       
9 (7) Petiole winged.   8 A. foliosa
+ Petiole wingless   (10)
       
10 (9) Leaf blade lanceolate, attenuate at base.   7 A. trinervis
+ Leaf blade ovate, rounded or cordate at base   (11)
       
11 (10) Petiole densely strigose or villous   (12)
+ Petiole glabrous   (13)
       
12 (11) Capitula usually in spikes; florets 3.   9 A. fragrans
+ Capitula in panicles; florets 3 or 4.   10 A. asaroides
       
13 (11) Leaf blade 2-3.5(-7) cm, usually caerulescent, caerulescent-purple, or purplish green abaxially.   11 A. gracilis
+ Leaf blade usually more than 4 cm, green abaxially   (14)
       
14 (13) Achenes glabrous or pilose above; leaf margin entire to shallowly denticulate.   12 A. kawakamii
+ Achenes pilose; leaf margin usually dentate   (15)
       
15 (14) Leaves aggregated near median part of stem.   13 A. macroclinidioides
+ Leaves above basal part of stem   (16)
       
16 (15) Petiole broadly winged, leaf blade margin obscurely denticulate.   14 A. smithii
+ Petiole wingless, leaf blade margin dentate.   15 A. aptera
       
17 (2) Leaf blade uniformly narrowed at base.   16 A. mairei
+ Leaf blade abruptly narrowed into petiole   (18)
       
18 (17) Leaves shortly petiolate (petioles 4-10 mm)   (19)
+ Leaves long petiolate (petioles more than 10 mm)   (20)
       
19 (18) Leaf blade lanceolate; achenes glabrous.   17 A. caesia
+ Leaf blade obovate to pandurate; achenes pilose.   18 A. spicata
       
20 (18) Petiole wholly winged (more than 1.5 mm wide at upper point) or winged in upper half   (21)
+ Petiole wingless (leaf blade cuneate in A. glabra, A. nervosa, and A. yunnanensis)   (26)
       
21 (20) Leaf base cordate.   19 A. bonatii
+ Leaf base rounded or attenuate   (22)
       
22 (21) Involucre 12-18 mm   (23)
+ Involucre less than 10 mm (occasionally equal to 10 mm)   (24)
       
23 (22) Anther appendages truncate; phyllaries and leaves pilose.   20 A. macrocephala
+ Anther appendages apiculate; phyllaries and leaves glabrous.   21 A. qianiana
       
24 (22) Leaf margin straight, leaf base rounded, petiole wholly winged.   22 A. latifolia
+ Leaf margin wavy, leaf base cuneate and commonly decurrent into petiole, petiole winged in upper half   (25)
       
25 (24) Achenes glabrous or apically pilose.   23 A. henryi
+ Achenes pilose.   24 A. reflexa
       
26 (20) Leaf margin dentate, blade obtuse to rounded at apex, discolorous, adaxial surface lanuginous and abaxial surface white tomentose.   25 A. crassifolia
+ Leaf margin entire to denticulate   (27)
       
27 (26) Leaf base cordate   (28)
+ Leaf base rounded or cuneate   (35)
       
28 (27) Capitula 2-flowered.   26 A. elegans
+ Capitula 3(or 4)-flowered   (29)
       
29 (28) Petiole lanuginous or glabrous   (30)
+ Petiole densely strigose   (32)
       
30 (29) Leaf blade 50-80 mm, apex acute to acuminate.   27 A. apteroides
+ Leaf blade less than 20 mm, apex acute or obtuse   (31)
       
31 (30) Leaf blade ovate, 7-16 × 5-10 mm; achenes glabrous.   28 A. nana
+ Leaf blade deltate, 10-15 × 10-15 mm; achenes pilose.   29 A. paucicapitata
       
32 (29) Anther appendages emarginate; achenes pilose above.   9 A. fragrans
+ Anther appendages apiculate; achenes pilose   (33)
       
33 (32) Phyllaries apically pilose.   30 A. cavaleriei
+ Phyllaries glabrous   (34)
       
34 (33) Leaf blade ± concolorous.   31 A. ramosa
+ Leaf blade abaxially whitish green, veins red.   32 A. rubrinervis
       
35 (27) Petiole densely strigose   (36)
+ Petiole lanuginous or glabrous   (38)
       
36 (35) Leaf blade elliptic.   33 A. angustata
+ Leaf blade oblong to elliptic   (37)
       
37 (36) Leaf blade 8-12 cm.   34 A. pingbianensis
+ Leaf blade 3-7.5 cm.   35 A. fulvipes
       
38 (35) Leaf blade elliptic, slightly fleshy, glabrous to subglabrous   (39)
+ Leaf blade ovate   (40)
       
39 (38) Leaf blade broadly elliptic, (2-)3.5-10 mm wide, apex acute-acuminate.   36 A. glabra
+ Leaf blade linear-elliptic, 1-2.5 mm wide, apex acute.   37 A. nervosa
       
40 (38) Achenes apically pilose.   38 A. parvifolia
+ Achenes pilose   (41)
       
41 (40) Leaf surface hairs T-shaped.   39 A. yunnanensis
+ Leaf surface hairs straight.   40 A. chapaensis

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