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6. Ajuga Linnaeus, Sp. Pl. 2: 561. 1753.
筋骨草属 jin gu cao shu
Plants annual, biennial or perennial, herbaceous, rarely shrubs. Leaves simple; leaf blade papery, margin dentate to incised, rarely subentire. Verticillasters 2- to many flowered, in false spikes; floral leaves similar to stem leaves or gradually reduced to bracts, rarely dissimilar, larger than stem leaves. Flowers subsessile. Calyx ovoid to globose, campanulate to funnelform, 10-veined, sometimes with inconspicuous accessory veins; teeth 5, slightly irregular. Corolla purple to blue, rarely yellow or white, 2-lipped, often persistent in fruit; tube straight to slightly curved, base slightly bent/swollen; throat slightly dilated, villous annulate, rarely glabrous inside; upper lip straight, entire to 2-lobed; lower lip elongate, 3-lobed, with middle lobe obcordate to nearly flabellate and lateral lobes oblong. Stamens 4, didynamous, exserted from upper lip, involute in bud, anterior 2 longer; filaments straight to slightly curved; anther cells 2, apically confluent. Style subequally 2-cleft, lobes subulate. Nutlets obovoid, triquetrous, netted on back, lateral-ventral side with an areole 1/2-2/3 its length, with an elaiosome.
About 40-50 species: Asia, Europe, especially in the Near East; 18 species in China.
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1 |
Verticillasters 2(-4, rarely more)-flowered, widely spaced; floral leaves similar to stem leaves; fertile branches (only) erect to ascending. |
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(2) |
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Verticillasters 6- or more flowered, close together; floral leaves unlike stem leaves (except in A. linearifolia); plants erect, rarely trailing |
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(5) |
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2 (1) |
Base of corolla tube not villous annulate inside; leaf base cordate to subtruncate |
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1 Ajuga lobata |
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Base of corolla tube villous annulate inside; leaf base cuneate to subtruncate, decurrent. |
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(3) |
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3 (2) |
Leaves spatulate, 2-3 × 4-6 mm |
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2 Ajuga pygmaea |
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Leaves ovate to oblong, more than 10 mm wide. |
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(4) |
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4 (3) |
Leaves 3-5.3 × 2.5-3.8 cm, ovate to ovate-oblong, basally cuneate to subtruncate, slightly decurrent |
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3 Ajuga sciaphila |
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Leaves 2-3.4 × 1.2-2.3 cm, oblong to ovate-oblong, basally cuneate to decurrent |
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4 Ajuga nubigena |
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5 (1) |
Floral leaves longer than flowers, whitish yellow, white, or purple |
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5 Ajuga lupulina |
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Floral leaves equal to or slightly shorter than flowers, rarely longer, green or purplish |
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(6) |
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6 (5) |
Corolla tube (1.5–)2–3 cm, upper lip 2-lobed |
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6 Ajuga ovalifolia |
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Corolla tube less than 1.6 cm, rarely to 2 cm, upper lip shallowly 2-lobed |
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(7) |
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7 (6) |
Floral and stem leaves similar, of 1 color; stem leaves linear to linear-lanceolate |
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18 Ajuga linearifolia |
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Floral and stem leaves dissimilar; stem leaves of other shapes |
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(8) |
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8 (7) |
Corolla tube erect to slightly curved, not saccate or bent above annulus of hairs |
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(9) |
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Corolla tube slightly swollen, shallowly saccate to bent above annulus of hairs |
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(14) |
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9 (8) |
Calyx ciliate along veins and teeth margin, villous to puberulent apically, glabrous basally, or subglabrous throughout |
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(10) |
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Calyx strigose, villous, or lanate-villous throughout, especially on teeth |
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(12) |
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10 (9) |
Stems 25–40 cm tall, purplish red or greenish purple, glabrous, gray villous only on younger parts; leaves ovate-oblong to narrowly oblong, 4–7.5?×?3.2–4 cm, irregularly double dentate |
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7 Ajuga ciliata |
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Stems less than 20 cm tall, sometimes to 30 cm, not brightly colored, densely gray pubescent to gray villous; leaves ovate to lanceolate-oblong, less than 3.2 cm wide, margin repand |
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(11) |
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11 (10) |
Inflorescences 1–3 cm; floral leaves usually shorter than flowers; calyx white villous on teeth |
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9 Ajuga campylanthoides |
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Inflorescences ca. 6 cm; floral leaves much longer than flowers; calyx ciliate along veins and apical teeth margin |
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11 Ajuga forrestii |
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12 (9) |
Calyx strigose; corolla tube slightly curved; stamens nearly included; plants subprostrate, basally stoloniferous |
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10 Ajuga campylantha |
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Calyx villous or lanate-villous; corolla tube straight; stamens slightly exserted; plants erect or stoloniferous |
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(13) |
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13 (12) |
Stems erect, unbranched, 6–21 cm tall, densely gray lanate or silky |
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8 Ajuga multiflora |
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Plants with stolons and erect fertile stems ca. 10 cm tall, gray villous or lanate-villous |
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12 Ajuga bracteosa |
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14 (8) |
Calyx teeth lanceolate to lanceolate-triangular; plants spreading ascending or with stolons |
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13 Ajuga pantantha |
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Calyx teeth narrowly triangular to broadly ovate; plants erect, sometimes trailing-ascending |
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(15) |
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15 (14) |
Corolla 6–8(–9) mm; calyx teeth ovate to ovate-triangular; stamens exserted ca. 1/2 their length |
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(16) |
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Corolla more than 8 mm; calyx teeth narrowly or short triangular; stamens exserted 1/4–1/3 their length |
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(17) |
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16 (15) |
Corolla white, upper lip shorter than lateral lobes of lower lip; calyx teeth ovate-triangular |
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16 Ajuga dictyocarpa |
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Corolla blue, blue-purple, or purple, upper lip nearly as long as lateral lobes of lower lip; calyx teeth ovate to broadly ovate |
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17 Ajuga macrosperma |
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17 (15) |
Leaves spatulate to suboblong; plants prostrate, with basal leaves at anthesis |
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14 Ajuga decumbens |
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Leaves broadly oblong to obovate-elliptic; plants erect or rarely prostrate, branched at base, without basal leaves at anthesis |
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15 Ajuga nipponensis |
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List of lower taxa
Related Links (opens in a new window) |
Treatments in Other Floras @ www.efloras.org
Other Databases
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