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6. Ajuga Linnaeus, Sp. Pl.  2: 561.  1753.  
筋骨草属 jin gu cao shu 
 
 
 
 
Plants annual, biennial or perennial, herbaceous, rarely shrubs. Leaves simple; leaf blade papery, margin dentate to incised, rarely subentire. Verticillasters 2- to many flowered, in false spikes; floral leaves similar to stem leaves or gradually reduced to bracts, rarely dissimilar, larger than stem leaves. Flowers subsessile. Calyx ovoid to globose, campanulate to funnelform, 10-veined, sometimes with inconspicuous accessory veins; teeth 5, slightly irregular. Corolla purple to blue, rarely yellow or white, 2-lipped, often persistent in fruit; tube straight to slightly curved, base slightly bent/swollen; throat slightly dilated, villous annulate, rarely glabrous inside; upper lip straight, entire to 2-lobed; lower lip elongate, 3-lobed, with middle lobe obcordate to nearly flabellate and lateral lobes oblong. Stamens 4, didynamous, exserted from upper lip, involute in bud, anterior 2 longer; filaments straight to slightly curved; anther cells 2, apically confluent. Style subequally 2-cleft, lobes subulate. Nutlets obovoid, triquetrous, netted on back, lateral-ventral side with an areole 1/2-2/3 its length, with an elaiosome. 
 
 
 
 
About 40-50 species: Asia, Europe, especially in the Near East; 18 species in China. 
 
 
 
  
                        
                         
		
                          
                          
			
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  | 1 | 
  Verticillasters 2(-4, rarely more)-flowered, widely spaced; floral leaves similar to stem leaves; fertile branches (only) erect to ascending. | 
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  (2) | 
 
  | + | 
  Verticillasters 6- or more flowered, close together; floral leaves unlike stem leaves (except in A. linearifolia); plants erect, rarely trailing | 
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  (5) | 
 
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  | 2 (1) | 
  Base of corolla tube not villous annulate inside; leaf base cordate to subtruncate | 
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  1 Ajuga lobata | 
 
  | + | 
  Base of corolla tube villous annulate inside; leaf base cuneate to subtruncate, decurrent. | 
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  (3) | 
 
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  | 3 (2) | 
  Leaves spatulate, 2-3 × 4-6 mm | 
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  2 Ajuga pygmaea | 
 
  | + | 
  Leaves ovate to oblong, more than 10 mm wide. | 
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  (4) | 
 
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  | 4 (3) | 
  Leaves 3-5.3 × 2.5-3.8 cm, ovate to ovate-oblong, basally cuneate to subtruncate, slightly decurrent | 
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  3 Ajuga sciaphila | 
 
  | + | 
  Leaves 2-3.4 × 1.2-2.3 cm, oblong to ovate-oblong, basally cuneate to decurrent | 
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  4 Ajuga nubigena | 
 
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  | 5 (1) | 
  Floral leaves longer than flowers, whitish yellow, white, or purple | 
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  5 Ajuga lupulina | 
 
  | + | 
  Floral leaves equal to or slightly shorter than flowers, rarely longer, green or purplish | 
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  (6) | 
 
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  | 6 (5) | 
  Corolla tube (1.5–)2–3 cm, upper lip 2-lobed | 
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  6 Ajuga ovalifolia | 
 
  | + | 
  Corolla tube less than 1.6 cm, rarely to 2 cm, upper lip shallowly 2-lobed | 
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  (7) | 
 
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  | 7 (6) | 
  Floral and stem leaves similar, of 1 color; stem leaves linear to linear-lanceolate | 
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  18 Ajuga linearifolia | 
 
  | + | 
  Floral and stem leaves dissimilar; stem leaves of other shapes | 
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  (8) | 
 
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  | 8 (7) | 
  Corolla tube erect to slightly curved, not saccate or bent above annulus of hairs | 
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  (9) | 
 
  | + | 
  Corolla tube slightly swollen, shallowly saccate to bent above annulus of hairs | 
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  (14) | 
 
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  | 9 (8) | 
  Calyx ciliate along veins and teeth margin, villous to puberulent apically, glabrous basally, or subglabrous throughout | 
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  (10) | 
 
  | + | 
  Calyx strigose, villous, or lanate-villous throughout, especially on teeth | 
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  (12) | 
 
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  | 10 (9) | 
  Stems 25–40 cm tall, purplish red or greenish purple, glabrous, gray villous only on younger parts; leaves ovate-oblong to narrowly oblong, 4–7.5?×?3.2–4 cm, irregularly double dentate | 
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  7 Ajuga ciliata | 
 
  | + | 
  Stems less than 20 cm tall, sometimes to 30 cm, not brightly colored, densely gray pubescent to gray villous; leaves ovate to lanceolate-oblong, less than 3.2 cm wide, margin repand | 
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  (11) | 
 
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  | 11 (10) | 
  Inflorescences 1–3 cm; floral leaves usually shorter than flowers; calyx white villous on teeth | 
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  9 Ajuga campylanthoides | 
 
  | + | 
  Inflorescences ca. 6 cm; floral leaves much longer than flowers; calyx ciliate along veins and apical teeth margin | 
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  11 Ajuga forrestii | 
 
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  | 12 (9) | 
  Calyx strigose; corolla tube slightly curved; stamens nearly included; plants subprostrate, basally stoloniferous | 
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  10 Ajuga campylantha | 
 
  | + | 
  Calyx villous or lanate-villous; corolla tube straight; stamens slightly exserted; plants erect or stoloniferous | 
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  (13) | 
 
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  | 13 (12) | 
  Stems erect, unbranched, 6–21 cm tall, densely gray lanate or silky | 
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  8 Ajuga multiflora | 
 
  | + | 
  Plants with stolons and erect fertile stems ca. 10 cm tall, gray villous or lanate-villous | 
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  12 Ajuga bracteosa | 
 
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  | 14 (8) | 
  Calyx teeth lanceolate to lanceolate-triangular; plants spreading ascending or with stolons | 
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  13 Ajuga pantantha | 
 
  | + | 
  Calyx teeth narrowly triangular to broadly ovate; plants erect, sometimes trailing-ascending | 
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  (15) | 
 
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  | 15 (14) | 
  Corolla 6–8(–9) mm; calyx teeth ovate to ovate-triangular; stamens exserted ca. 1/2 their length | 
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  (16) | 
 
  | + | 
  Corolla more than 8 mm; calyx teeth narrowly or short triangular; stamens exserted 1/4–1/3 their length | 
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  (17) | 
 
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  | 16 (15) | 
  Corolla white, upper lip shorter than lateral lobes of lower lip; calyx teeth ovate-triangular | 
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  16 Ajuga dictyocarpa | 
 
  | + | 
  Corolla blue, blue-purple, or purple, upper lip nearly as long as lateral lobes of lower lip; calyx teeth ovate to broadly ovate | 
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  17 Ajuga macrosperma | 
 
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  | 17 (15) | 
  Leaves spatulate to suboblong; plants prostrate, with basal leaves at anthesis | 
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  14 Ajuga decumbens | 
 
  | + | 
  Leaves broadly oblong to obovate-elliptic; plants erect or rarely prostrate, branched at base, without basal leaves at anthesis | 
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  15 Ajuga nipponensis | 
 
 
 
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                          List of lower taxa 
 
                        
                          
                        
	
                        
                      
 
                      
	 
                      
                         
		
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