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36. Amitostigma Schlechter, Repert. Spec. Nov. Regni Veg. Beih. 4: 91. 1919.

无柱兰属 wu zhu lan shu

Authors: Xinqi Chen, Stephan W. Gale & Phillip J. Cribb

Mitostigma Blume, Mus. Bot. 2: 189. 1856, not Decaisne (1844).

Herbs, terrestrial, slender. Tubers subglobose to ovoid, undivided, fleshy, neck with several slender roots. Stem erect, 1- or occasionally 2-leaved, glabrous or occasionally pubescent. Leaves basal or cauline, usually green and unspotted, rarely with purple-brown blotches, oblong, lanceolate, elliptic, or ovate, base contracted into amplexicaul sheath. Inflorescence a terminal raceme, glabrous; peduncle elongate, with 1 or more sterile bracts; rachis 1- to several flowered. Flowers often secund, resupinate, white, pink, mauve, or purple, rarely yellow; ovary usually straight, twisted, shortly pedicellate, cylindric-fusiform, glabrous. Sepals free, oblong to ovate, 1-veined. Petals free, often connivent with dorsal sepal and forming a hood, usually wider than sepals; lip spreading, spurred at base, 3-lobed; disk often papillose; mid-lobe often divided, forming 2 lobules; spur cylindric to narrowly conic, short to elongate. Column very short; anther usually erect, oblong-ellipsoid to obovoid, 2-locular, locules parallel or slightly divergent; pollinia 2, oblong, sectile, granular-farinaceous; caudicles straplike; viscidia naked, oblong-ovate; rostellum rather small, 3-lobed, base with ± ascending arms on both sides, margin adnate to column; stigma ventral, 3-lobed, lobes confluent, relatively large, flat to somewhat raised; auricles 2, large, stout, placed laterally at base of anther. Capsule suberect.

About 30 species: E Asia and adjacent areas; 22 species (21 endemic) in China.

The generic affinity of several of the entities presently referred to Amitostigma remains questionable and in need of re-examination; many of the endemic Chinese taxa are poorly known.


1 Plants 2-leaved (A. alpestre rarely 1-leaved)   (2)
+ Plants 1-leaved   (7)
       
2 (1) Leaves cauline, widely spaced   (3)
+ Leaves basal, subopposite   (4)
       
3 (2) Lip obovate, 3-lobed near middle, 4.5-5.5 mm wide, longer than broad; lateral lobes ovate-oblong, ca. 0.5 mm wide; spur 4-4.5 mm.   17 A. alpestre
+ Lip orbicular to transversely elliptic, 3-lobed toward apex, ca. 8.5 mm wide, broader than long; lateral lobes quadrate, ca. 2.5 mm wide; spur 5-7 mm.   18 A. yuanum
       
4 (2) Rachis 1-flowered.   19 A. papilionaceum
+ Rachis 3- to more than 10-flowered   (5)
       
5 (4) Leaves usually appressed to substratum; petals ca. 5 × 3 mm; lip ca. 9 mm.   22 A. physoceras
+ Leaves erect to porrect; petals 3-3.5 × 1.5-2 mm; lip 4.5-6 mm   (6)
       
6 (5) Lip 3-lobed below middle; lateral lobes linear.   20 A. bifoliatum
+ Lip 3-lobed above middle; lateral lobes ovate.   21 A. wenshanense
       
7 (1) Leaf appressed to substratum, adaxially with purplish brown blotches.   1 A. hemipilioides
+ Leaf erect or porrect, uniformly green   (8)
       
8 (7) Rachis 1- or rarely 2-flowered   (9)
+ Rachis usually 3- or more flowered (A. faberi rarely 1- or 2-flowered)   (15)
       
9 (8) Spur 8-17 mm, ca. as long as to exceeding ovary   (10)
+ Spur 2-4 mm, usually much shorter than ovary   (12)
       
10 (9) Lip 3-lobed toward apex, 11-15 × 13-19 mm; lateral lobes broadly oblong-ovate, ca. 5 mm wide.   6 A. pinguicula
+ Lip 3-lobed below middle, 8-10 × 6-10 mm; lateral lobes linear-oblong or ovate-triangular, 1-3 mm wide   (11)
       
11 (10) Dorsal sepal 4-5 mm; spur 11-12 mm.   7 A. dolichocentrum
+ Dorsal sepal ca. 7 mm; spur 8-9 mm.   8 A. tibeticum
       
12 (9) Stem densely pubescent around leaf base; floral bract much shorter than ovary; lip 5-6 mm.   2 A. amplexifolium
+ Stem glabrous; floral bract nearly as long as to exceeding ovary; lip 7-15 mm   (13)
       
13 (12) Flower yellow.   3 A. simplex
+ Flower pale purple, pink, or white   (14)
       
14 (13) Leaf 0.1-0.4 cm wide; petals broadly ovate, 4.5-5.5 mm wide.   4 A. farreri
+ Leaf 0.6-1 cm wide; petals ovate, 1.6-2 mm wide.   5 A. monanthum
       
15 (8) Mid-lobe of lip 4-7 mm wide, nearly as broad as to broader than long   (16)
+ Mid-lobe of lip 0.5-3.5 mm wide, much longer than broad   (18)
       
16 (15) Ovary and pedicel 15-25 mm; dorsal sepal ca. 7 mm; lip ca. 13 mm.   16 A. gonggashanicum
+ Ovary and pedicel 6-10 mm; dorsal sepal 2.5-3.5 mm; lip 5-7.5 mm   (17)
       
17 (16) Peduncle ebracteate; spur 2-3 mm.   14 A. basifoliatum
+ Peduncle with 1 sterile bract; spur 5-6 mm.   15 A. faberi
       
18 (15) Mid-lobe of lip ca. 3.5 mm wide, deeply divided above middle to form 2 distinct lobules.   9 A. parceflorum
+ Mid-lobe of lip 0.5-2 mm wide, apex truncate, obtuse-rounded, occasionally emarginate or shallowly notched   (19)
       
19 (18) Lateral lobes of lip oblong-obovate, 1.6-2 mm wide; spur 3.5-6 mm.   10 A. tetralobum
+ Lateral lobes of lip lanceolate, linear-oblong, ovate-oblong, or triangular, 0.3-0.8 mm wide; spur 1-2.5(-3.5) mm   (20)
       
20 (19) Peduncle ebracteate; flowers white; mid-lobe of lip ca. 1.6 mm.   11 A. capitatum
+ Peduncle with 1 or 2 sterile bracts; flowers pink, rose-pink, purplish red, or purple; mid-lobe of lip 2-4 mm   (21)
       
21 (20) Plants 7-25 cm tall; leaf usually basal; lip 3-lobed above middle.   12 A. gracile
+ Plants 24-36 cm tall; leaf cauline; lip 3-lobed below middle.   13 A. trifurcatum

  • List of lower taxa


     

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