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36. Amitostigma Schlechter, Repert. Spec. Nov. Regni Veg. Beih. 4: 91. 1919.
无柱兰属 wu zhu lan shu
Authors: Xinqi Chen, Stephan W. Gale & Phillip J. Cribb
Mitostigma Blume, Mus. Bot. 2: 189. 1856, not Decaisne (1844).
Herbs, terrestrial, slender. Tubers subglobose to ovoid, undivided, fleshy, neck with several slender roots. Stem erect, 1- or occasionally 2-leaved, glabrous or occasionally pubescent. Leaves basal or cauline, usually green and unspotted, rarely with purple-brown blotches, oblong, lanceolate, elliptic, or ovate, base contracted into amplexicaul sheath. Inflorescence a terminal raceme, glabrous; peduncle elongate, with 1 or more sterile bracts; rachis 1- to several flowered. Flowers often secund, resupinate, white, pink, mauve, or purple, rarely yellow; ovary usually straight, twisted, shortly pedicellate, cylindric-fusiform, glabrous. Sepals free, oblong to ovate, 1-veined. Petals free, often connivent with dorsal sepal and forming a hood, usually wider than sepals; lip spreading, spurred at base, 3-lobed; disk often papillose; mid-lobe often divided, forming 2 lobules; spur cylindric to narrowly conic, short to elongate. Column very short; anther usually erect, oblong-ellipsoid to obovoid, 2-locular, locules parallel or slightly divergent; pollinia 2, oblong, sectile, granular-farinaceous; caudicles straplike; viscidia naked, oblong-ovate; rostellum rather small, 3-lobed, base with ± ascending arms on both sides, margin adnate to column; stigma ventral, 3-lobed, lobes confluent, relatively large, flat to somewhat raised; auricles 2, large, stout, placed laterally at base of anther. Capsule suberect.
About 30 species: E Asia and adjacent areas; 22 species (21 endemic) in China.
The generic affinity of several of the entities presently referred to Amitostigma remains questionable and in need of re-examination; many of the endemic Chinese taxa are poorly known.
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1 |
Plants 2-leaved (A. alpestre rarely 1-leaved) |
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(2) |
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Plants 1-leaved |
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(7) |
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2 (1) |
Leaves cauline, widely spaced |
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(3) |
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Leaves basal, subopposite |
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(4) |
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3 (2) |
Lip obovate, 3-lobed near middle, 4.5-5.5 mm wide, longer than broad; lateral lobes ovate-oblong, ca. 0.5 mm wide; spur 4-4.5 mm. |
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17 A. alpestre |
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Lip orbicular to transversely elliptic, 3-lobed toward apex, ca. 8.5 mm wide, broader than long; lateral lobes quadrate, ca. 2.5 mm wide; spur 5-7 mm. |
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18 A. yuanum |
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4 (2) |
Rachis 1-flowered. |
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19 A. papilionaceum |
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Rachis 3- to more than 10-flowered |
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(5) |
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5 (4) |
Leaves usually appressed to substratum; petals ca. 5 × 3 mm; lip ca. 9 mm. |
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22 A. physoceras |
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Leaves erect to porrect; petals 3-3.5 × 1.5-2 mm; lip 4.5-6 mm |
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(6) |
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6 (5) |
Lip 3-lobed below middle; lateral lobes linear. |
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20 A. bifoliatum |
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Lip 3-lobed above middle; lateral lobes ovate. |
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21 A. wenshanense |
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7 (1) |
Leaf appressed to substratum, adaxially with purplish brown blotches. |
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1 A. hemipilioides |
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Leaf erect or porrect, uniformly green |
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(8) |
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8 (7) |
Rachis 1- or rarely 2-flowered |
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(9) |
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Rachis usually 3- or more flowered (A. faberi rarely 1- or 2-flowered) |
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(15) |
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9 (8) |
Spur 8-17 mm, ca. as long as to exceeding ovary |
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(10) |
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Spur 2-4 mm, usually much shorter than ovary |
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(12) |
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10 (9) |
Lip 3-lobed toward apex, 11-15 × 13-19 mm; lateral lobes broadly oblong-ovate, ca. 5 mm wide. |
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6 A. pinguicula |
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Lip 3-lobed below middle, 8-10 × 6-10 mm; lateral lobes linear-oblong or ovate-triangular, 1-3 mm wide |
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(11) |
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11 (10) |
Dorsal sepal 4-5 mm; spur 11-12 mm. |
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7 A. dolichocentrum |
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Dorsal sepal ca. 7 mm; spur 8-9 mm. |
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8 A. tibeticum |
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12 (9) |
Stem densely pubescent around leaf base; floral bract much shorter than ovary; lip 5-6 mm. |
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2 A. amplexifolium |
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Stem glabrous; floral bract nearly as long as to exceeding ovary; lip 7-15 mm |
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(13) |
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13 (12) |
Flower yellow. |
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3 A. simplex |
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Flower pale purple, pink, or white |
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(14) |
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14 (13) |
Leaf 0.1-0.4 cm wide; petals broadly ovate, 4.5-5.5 mm wide. |
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4 A. farreri |
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Leaf 0.6-1 cm wide; petals ovate, 1.6-2 mm wide. |
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5 A. monanthum |
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15 (8) |
Mid-lobe of lip 4-7 mm wide, nearly as broad as to broader than long |
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(16) |
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Mid-lobe of lip 0.5-3.5 mm wide, much longer than broad |
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(18) |
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16 (15) |
Ovary and pedicel 15-25 mm; dorsal sepal ca. 7 mm; lip ca. 13 mm. |
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16 A. gonggashanicum |
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Ovary and pedicel 6-10 mm; dorsal sepal 2.5-3.5 mm; lip 5-7.5 mm |
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(17) |
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17 (16) |
Peduncle ebracteate; spur 2-3 mm. |
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14 A. basifoliatum |
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Peduncle with 1 sterile bract; spur 5-6 mm. |
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15 A. faberi |
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18 (15) |
Mid-lobe of lip ca. 3.5 mm wide, deeply divided above middle to form 2 distinct lobules. |
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9 A. parceflorum |
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Mid-lobe of lip 0.5-2 mm wide, apex truncate, obtuse-rounded, occasionally emarginate or shallowly notched |
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(19) |
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19 (18) |
Lateral lobes of lip oblong-obovate, 1.6-2 mm wide; spur 3.5-6 mm. |
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10 A. tetralobum |
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Lateral lobes of lip lanceolate, linear-oblong, ovate-oblong, or triangular, 0.3-0.8 mm wide; spur 1-2.5(-3.5) mm |
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(20) |
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20 (19) |
Peduncle ebracteate; flowers white; mid-lobe of lip ca. 1.6 mm. |
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11 A. capitatum |
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Peduncle with 1 or 2 sterile bracts; flowers pink, rose-pink, purplish red, or purple; mid-lobe of lip 2-4 mm |
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(21) |
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21 (20) |
Plants 7-25 cm tall; leaf usually basal; lip 3-lobed above middle. |
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12 A. gracile |
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Plants 24-36 cm tall; leaf cauline; lip 3-lobed below middle. |
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13 A. trifurcatum |
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List of lower taxa
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