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Crassulaceae Candolle
景天科 jing tian ke
Authors: Kunjun Fu, Hideaki Ohba & Michael G. Gilbert
Herbs, subshrubs, or shrubs. Stems mostly fleshy. Leaves alternate, opposite, or verticillate, usually simple; stipules absent; leaf blade entire or slightly incised, rarely lobed or imparipinnate. Inflorescences terminal or axillary, cymose, corymbiform, spiculate, racemose, paniculate, or sometimes reduced to a solitary flower. Flowers usually bisexual, sometimes unisexual in Rhodiola (when plants dioecious or rarely gynodioecious), actinomorphic, (3 or)4-6(-30)-merous. Sepals almost free or basally connate, persistent. Petals free or connate. Stamens as many as petals in 1 series or 2 × as many in 2 series. Nectar scales at or near base of carpels. Follicles sometimes fewer than sepals, free or basally connate, erect or spreading, membranous or leathery, 1- to many seeded. Seeds small; endosperm scanty or not developed.
About 35 genera and over 1500 species: Africa, America, Asia, Europe; 13 genera (two endemic, one introduced) and 233 species (129 endemic, one introduced) in China.
Some species of Crassulaceae are cultivated as ornamentals and/or used medicinally. Fu Shu-hsia & Fu Kun-tsun. 1984. Crassulaceae. In: Fu Shu-hsia & Fu Kun-tsun, eds., Fl. Reipubl. Popularis Sin. 34(1): 31-220.
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| 1 |
Stamens in 1 series, usually as many as petals; flowers always bisexual. |
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(2) |
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Stamens usually in 2 series, 2 × as many as petals; flowers sometimes unisexual and male flowers without stamens. |
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(5) |
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| 2 (1) |
Leaves always opposite, joined to form a basal sheath; inflorescences axillary, often shorter than subtending leaf; plants not developing enlarged rootstock |
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1 Tillaea |
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Leaves alternate, occasionally opposite proximally; inflorescence terminal, often very large; plants sometimes developing enlarged, perennial rootstock. |
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(3) |
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| 3 (2) |
Inflorescence with a distinct, elongated main axis forming a raceme or thyrse much longer than broad; carpels narrowed at base, free; basal leaves often densely hairy |
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4 Kungia |
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Inflorescence cymose with no distinct main axis, often broader than long; carpels broad at base, often ± connate; basal leaves often glabrous. |
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(4) |
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| 4 (3) |
Plants with conspicuous basal leaf rosettes; corolla campanulate, petals initially erect then spreading above middle, ± S-shaped in longitudinal section |
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8 Sinocrassula |
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Plants without basal leaf rosettes, often ephemeral; petals rotate |
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12 Sedum |
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| 5 (1) |
Flowers 4-merous and with prominent, persistent corolla tube completely enclosing follicles; leaves opposite. |
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(6) |
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Flowers 5- or 6(-12)-merous, rarely 3- or 4-merous (Rhodiola) and then petals free or joined at base only and not enclosing follicles; leaves alternate, opposite, or in a rosette. |
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(7) |
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| 6 (5) |
Filaments inserted basally on corolla tube; flowers usually pendulous |
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2 Bryophyllum |
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Filaments inserted near or above middle of corolla tube; flowers erect |
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3 Kalanchoe |
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| 7 (5) |
Carpels stipitate to substipitate or basally constricted or conspicuously attenuate, free. |
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(8) |
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Carpels sessile, base not constricted (slightly attenuate in some Rhodiola species) and usually connate (free in Sedum chuhsingense, S. semilunatum, and sometimes in S. przewalskii). |
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(9) |
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| 8 (7) |
Plants perennial, usually with several to many flowering stems; basal leaves not in a conspicuous rosette; inflorescence formed from a terminal group of cymes, usually broader than long, sometimes with aditional cymes in axils of distal leaves |
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6 Hylotelephium |
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Plants monocarpic, with a single flowering stem; basal leaves in a somewhat conspicuous rosette, dying when plant flowers; inflorescence with an elongated main axis and numerous lateral cymules, sometimes a true raceme with cymules reduced to single flowers |
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5 Orostachys |
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| 9 (7) |
Stems dimorphic with usually very stout caudex or rhizome, usually with brown or blackish, membranous, scalelike leaves, sharply differentiated from much more slender, erect or ascending, leafy flowering stems. |
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(10) |
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Stems not sharply dimorphic, proximally with well developed leaves, rarely plants stoloniferous. |
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(11) |
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| 10 (9) |
Flowers bisexual; petals basally connate |
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7 Pseudosedum |
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Flowers unisexual or bisexual; petals free or nearly so |
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13 Rhodiola |
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| 11 (9) |
Leaf rosette absent at anthesis; inflorescence terminal. |
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(12) |
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Leaf rosette conspicuous at anthesis; inflorescence axillary. |
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(13) |
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| 12 (11) |
Leaves flattened, margin serrate or crenate; seed testa longitudinally costate or subsmooth |
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11 Phedimus |
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Leaves terete or semiterete in cross section, margin entire (4-8-dentate in Sedum rosthornianum and remotely dentate in S. engleri var. dentatum); seed testa reticulate or papillate-reticulate |
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12 Sedum |
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| 13 (11) |
Petals yellow; basal leaf rosette almost as wide as inflorescences are long; plants glabrous throughout |
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10 Ohbaea |
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Petals white, pink, and/or red, (rarely pale yellow outside China); plants hairy or glabrous. |
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(14) |
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| 14 (13) |
Plants glandular hairy or, if glabrous, flowers 6-8-merous; petals connate at base |
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9 Rosularia |
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Plants glabrous; flowers (3-)5-merous; petals free or almost so |
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13 Rhodiola |
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List of lower taxa
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