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FOC | Family List | FOC Vol. 6 | Ranunculaceae

35. Batrachium (de Candolle) Gray, Nat. Arr. Brit. Pl. 2: 720. 1821.

水毛茛属 shui mao gen shu

Wang Wencai; Michio Tamura

Ranunculus sect. Batrachium de Candolle, Syst. Nat. 1: 233. 1817.

Herbs perennial or annual, aquatic or semiterrestrial, with stems and leaves often submersed. Leaves alternate, subsessile to long petiolate, leaf blade usually submersed, 2--5 × dissected, ultimate segments filiform or narrowly linear, sometimes floating and then blade 3-lobed, not finely dissected. Inflorescence a solitary, leaf-opposed flower. Flowers bisexual, actinomorphic. Receptacle conical or subulate. Sepals (4 or)5, caducous. Petals (4 or)5, usually white, with yellow base, rarely entirely yellow, obovate, base shortly clawed, with a nectary pit above claw, apex rounded. Stamens few or numerous; anthers latrorse. Carpels numerous; ovule 1 per carpel. Style with adaxial stigmatic surface. Aggregate fruit ovoid or globose; achenes obovoid, slightly bilaterally compressed, transversely rugose.

About 20 species: S Africa, Asia, SE Australia, Europe, North and South America; eight species (one endemic) in China.

The separation of Batrachium from Ranunculus is not clear-cut and many botanists prefer to include the former genus within the latter. All species are aquatic or grow on wet mud.

1 Leaves dimorphic: submerged leaves 3--4 × sect with filiform segments, floating leaves ca. 2 × sect with linear- lanceolate segments   1 B. pekinense
+ Leaves all similar, usually submerged, with filiform segments.   (2)
2 (1) Receptacle glabrous   2 B. kauffmanii
+ Receptacle puberulent.   (3)
3 (2) Receptacle subulate in fruit   8 B. rionii
+ Receptacle conical in fruit.   (4)
4 (3) Ultimate leaf segments soft, collapsing out of water   4 B. bungei
+ Ultimate leaf segments rigid, ± divaricate out of water.   (5)
5 (4) Leaf blade orbicular in outline   7 B. foeniculaceum
+ Leaf blade flabellate or semiorbicular in outline.   (6)
6 (5) Herbs terrestrial or growing in shallow water, not submersed, ± erect; stems 1.5--14 cm   6 B. eradicatum
+ Herbs submerged; stems up to 40 cm.   (7)
7 (6) Stem and petiole sheath glabrous   3 B. divaricatum
+ Stem usually sparsely puberulent; petiole sheath abaxially strigose   5 B. trichophyllum

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