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2. Brandisia J. D. Hooker & Thomson, J. Linn. Soc., Bot. 8: 11. 1864.

来江藤属 lai jiang teng shu

Shrubs, erect or scandent, occasionally parasitic, usually stellate tomentose. Leaves opposite, rarely subopposite; petiole short; leaf blade margin entire or denticulate. Flowers axillary, solitary, paired, or in racemes. Bracteoles 2, inserted on pedicels. Calyx campanulate, rarely cylindric-ovate, outside stellate hairy, regular or somewhat 2-lipped; lobes 2 or 5. Corolla tube somewhat incurved; limb 2-lipped; lower lip as long as or shorter than upper lip, 3-lobed, patent; upper lip 2-lobed, concave. Stamens 4, didynamous, somewhat exserted or included, inserted at base of corolla tube; filaments glabrous; anther locules 2, orbicular, pilose; staminode absent. Ovary ovoid, hairy, 2-loculed; ovules numerous, linear-oblong. Style elongated; stigma simple. Capsule ovoid to ovoid-globose, loculicidal. Seeds linear; seed coat reticulate, membranous winged.

About 11 species: mountainous areas of subtropical E Asia; eight species in China.


1 Stellate hairs early glabrescent; corolla lower lip ca. 1/2 as long as upper lip, inside pubescent; inflorescences distinctly racemose.   (2)
+ Stellate hairs persisting (late glabrescent in B. glabrescens var. glabrescens); corolla lower lip as long as or slightly shorter than upper lip; inflorescences solitary or paired flowers, rarely subracemose.   (3)
       
2 (1) Racemes mostly borne on lateral branches, to 20 cm or longer; bracts large; leaf blade ovate-orbicular to ovate-lanceolate, margin serrulate, apex acute   7 Brandisia racemosa
+ Racemes borne on main stem, 5-6 cm; bracts small; leaf blade narrowly lanceolate, margin entire, apex narrowly acuminate   8 Brandisia cauliflora
       
3 (1) Calyx obscurely veined; corolla more than 3 X as long as calyx   6 Brandisia rosea
+ Calyx conspicuously veined; corolla less than 3 X as long as calyx.   (4)
       
4 (3) Calyx lobes irregular, 2-lipped   5 Brandisia kwangsiensis
+ Calyx lobes regular, not 2-lipped.   (5)
       
5 (4) Calyx tubular-ovoid, length greater than width, apex not gaping   4 Brandisia glabrescens
+ Calyx broadly campanulate, length and width equal, apex gaping.   (6)
       
6 (5) Calyx tube 5-veined   3 Brandisia discolor
+ Calyx tube 10-veined.   (7)
       
7 (6) Tomentum dull yellow; leaf blade ovate-lanceolate; calyx lobe length equal or less than width; corolla orange-red   1 Brandisia hancei
+ Tomentum gray-brown; leaf blade ovate, rarely ovate-oblong; calyx lobe length more than width; corolla yellow   2 Brandisia swinglei

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