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3. Broussonetia L’Héritier ex Ventenat, Tabl. Règn. Vég. 3: 547. 1799.
[nom. cons., not Ortega (1798).]
构属 gou shu
Allaeanthus Thwaites; Smithiodendron Hu.
Trees, shrubs, or climbers, deciduous, with latex; monoecious or dioecious. Winter buds small. Stipules free, lateral, ovate-lanceolate, caducous. Leaves alternate, spirally arranged or distichous; leaf blade simple to palmately lobed, margin toothed; primary veins 3-5 and plinerved, secondary veins pinnate. Male inflorescences axillary, spicate or ± capitate, many-flowered. Female inflorescences densely capitate to ± elongated; bracts clavate, persistent. Male flowers: calyx (3 or) 4-lobed, valvate; stamens inflexed in bud. Female flowers: calyx lobes connate into a tube, apically entire or 3- or 4-lobed, persistent; ovary enclosed, stipitate; style lateral; stigma linear, often with vestigial second lobe at base. Fruit densely aggregated into globose syncarp. Syncarp globose; pericarp membranous, enlarged; exocarp fleshy. Cotyledons orbicular, flat, or conduplicate; embryo curved.
Four species: E Asia, Pacific Islands; four species in China.
The bark fibers are used for making paper, the wood is used for furniture, and the leaves and fruit are used medicinally.
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1 |
Flowers produced on leafless stems; leaves distichous; leaf blade glabrous or nearly so; endocarp smooth. |
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4 B. kurzii |
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Flowers always produced on leafy stems; leaves spirally arranged; leaf blade adaxially scabrous or scabridulous; endocarp minutely tuberculate. |
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(2) |
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2 (1) |
Stipules 15-20 × 8-10 mm; petiole 2.3-8 cm; leaf blade abaxially densely pubescent between pilose veins; syncarp 1.5-3 cm in diam., mostly pubescent with scattered stout, ± barbed hairs. |
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1 B. papyrifera |
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Stipules 3-5 × 0.5-1 mm; petiole 0.8-1 cm; leaf blade abaxially glabrous or sparsely pubescent, mainly on veins; syncarp 0.8-1 cm in diam., with stellate clusters of stout strongly barbed hairs. |
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(3) |
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3 (2) |
Plant monoecious, freestanding; male inflorescences globose, 0.8-1 cm. |
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2 B. kazinoki |
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Plant dioecious, climbing; male inflorescences spicate, 1.5-2.5 cm. |
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3 B. kaempferi var. australis |
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Lower Taxa
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