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FOC | Family List | FOC Vol. 12 | Theaceae

10. Cleyera Thunberg, Nov. Gen. Pl. 68. 1783.

红淡比属 hong dan bi shu

Sakakia Nakai; Tristylium Turczaninow.

Shrubs or trees, evergreen. Leaves petiolate; leaf blade margin entire or serrulate. Flowers bisexual, axillary, solitary or several in a cluster. Pedicel 1 cm or more, apically thickened; bracteoles 2, caducous, small, inserted near pedicel apex. Sepals 5, imbricate, unequal, basally slightly connate; outer sepals persistent, small. Petals 5, imbricate, basally connate. Stamens 25-30; filaments distinct, glabrous; anthers basifixed, 2-loculed, longitudinally dehiscent, with filiform trichomes, connective apiculate. Ovary usually glabrous, 2- or 3-loculed with 8-16 ovules per locule, placentation axile; style 1, persistent, slender, elongated, apically 2- or 3-lobed. Fruit baccate, ovoid to oblate, with several seeds per locule. Seeds blackish brown, reniform-globose to compressed globose, foveolate, shiny, glabrous; endosperm sparse; embryo curved.

About 24 species: China, N India, Japan, Korea, Myanmar, Nepal, N Vietnam; tropical America; nine species (seven endemic) in China.

Unlike Cleyera japonica, the Thunberg description of the genus Cleyera did not include elements of a mixed collection with Ternstroemia.

1 Fruit long ovoid, oblong, obconic, or ovoid-ellipsoid   (2)
+ Fruit globose   (4)
2 (1) Leaf blade ovate to obovate-oblong, margin entire; sepals orbicular.   8 C. obovata
+ Leaf blade oblong-elliptic or rarely oblong-oblanceolate, margin serrulate, sparsely serrate, or rarely subentire; sepals suborbicular, triangular, or ovate-triangular   (3)
3 (2) Leaf blade abaxially reddish brown glandular punctate; sepals suborbicular.   7 C. obscurinervia
+ Leaf blade abaxially not punctate; sepals triangular to ovate-triangular.   9 C. longicarpa
4 (1) Leaf blade abaxially not punctate   (5)
+ Leaf blade abaxially densely or sparsely reddish or reddish brown glandular punctate   (7)
5 (4) Sepals ovate-triangular, apex acute with a black mucro; petiole 3-5 mm; pedicel 0.5-0.8 cm.   3 C. parvifolia
+ Sepals ovate to orbicular, apex rounded and without a mucro; petiole 7-10(-12) mm; pedicel 1-2(-3) cm   (6)
6 (5) Leaf blade margin entire, midvein adaxially slightly impressed.   1 C. japonica
+ Leaf blade margin conspicuously serrate, crenate serrate, or sparsely serrate, midvein adaxially raised.   2 C. lipingensis
7 (4) Leaf blade oblong, secondary veins 20-28 on each side of midvein; sepals oblong to ovate-oblong, thick.   4 C. pachyphylla
+ Leaf blade elliptic, oblong-elliptic, narrowly elliptic, or oblanceolate-elliptic, secondary veins 8-12 on each side of midvein; sepals ovate to orbicular, thin   (8)
8 (7) Leaf blade elliptic to oblong-elliptic, margin entire, midvein adaxially raised, secondary veins 12-14 on each side of midvein.   5 C. yangchunensis
+ Leaf blade narrowly elliptic to oblanceolate-elliptic, margin serrulate to sparsely serrate, midvein adaxially slightly impressed, secondary veins 8-12 on each side of midvein.   6 C. incornuta

  • List of lower taxa


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