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23. Cousinia Cassini in F. Cuvier, Dict. Sci. Nat. 47: 503. 1827.
刺头菊属 ci tou ju shu
Authors: Zhu Shi & Werner Greuter
Herbs, biennial or perennial. Leaves divided or undivided, margin spiny or spinulose. Capitula 1 to many, forming a racemose, paniculate, or corymbose synflorescence, or solitary, glabrous to cobwebby. Involucre spherical to cylindric. Phyllaries imbricate, leathery, ending in a spine. Stamen filaments glabrous; anther with ± lacerate basal appendages. Style branches short to long, apex acute. Achene obovoid to obconic, laterally compressed, with (2-)4 or 5 raised ribs, rugose, apex ± rimmed; rim sometimes denticulate. Pappus rarely absent, when present bristles in ca. 3 rows, outer ones much shorter than inner.
About 600 species: C, S, and SW Asia; 11 species (two endemic) in China.
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| 1 |
Stems winged |
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(2) |
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Stems unwinged |
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(5) |
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| 2 (1) |
Corolla yellow |
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(3) |
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Corolla red to purple |
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(4) |
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| 3 (2) |
Involucre 1.2-1.5 cm in diam.; leaves ± leathery. |
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1 C. dissecta |
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Involucre 0.6-0.9 cm in diam.; leaves thinly papery. |
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2 C. alata |
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| 4 (2) |
Involucre 1.2-1.5 cm in diam., glabrous; phyllaries green; leaves undivided. |
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3 C. leiocephala |
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Involucre 1.5-2 cm in diam., densely cobwebby; phyllaries grayish white; basal and lower cauline leaves pinnately lobed. |
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4 C. platylepis |
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| 5 (1) |
Corolla white or yellow |
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(6) |
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Corolla purplish red |
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(7) |
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| 6 (5) |
Involucre spherical to ovoid, 1.5-2 cm in diam.; corolla white. |
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5 C. affinis |
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Involucre narrowly ellipsoid to cylindric, 0.6-0.8 cm in diam.; corolla yellow. |
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6 C. polycephala |
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| 7 (5) |
Stems unbranched; capitulum solitary |
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(8) |
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Stems branched; capitula several |
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(9) |
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| 8 (7) |
Involucre 4-4.5 cm in diam.; leaf segments broadly ovate. |
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8 C. sclerolepis |
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Involucre 1.5-2 cm in diam.; leaf segments triangular to triangular-lanceolate. |
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9 C. caespitosa |
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| 9 (7) |
Leaves undivided. |
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7 C. lasiophylla |
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Basal and lower cauline leaves pectinately divided into needle-shaped segments |
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(10) |
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| 10 (9) |
Capitula 4 or 5; terminal spine of leaf segments 5-10 mm. |
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10 C. falconeri |
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Capitula numerous; terminal spine of leaf segments ca. 4 mm. |
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11 C. thomsonii |
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List of lower taxa
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Treatments in Other Floras @ www.efloras.org
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