All Floras      Advanced Search
FOC Vol. 20-21 Page 6, 198, 252, 264 Login | eFloras Home | Help
FOC | Family List | FOC Vol. 20-21 | Asteraceae

68. Crepidiastrum Nakai, Bot. Mag. (Tokyo). 34: 147. 1920.

假还阳参属 jia huan yang shen shu

Authors: Zhu Shi & Norbert Kilian

Crepidifolium Sennikov; Geblera Kitagawa (1937), not Fischer & C. A. Meyer (1835); Paraixeris Nakai.

Herbs, annual, biennial, or perennial, sometimes subshrubs, often rosulate, with a taproot. Stems usually leafy. Leaves undivided or pinnately lobed; stem leaves often clasping. Capitula with 5-20 florets. Involucres narrowly cylindric. Phyllaries with narrow scarious margin; outer phyllaries few, longest ca. 1/4(-1/2) as long as inner ones; inner phyllaries 5 or 8, linear-lanceolate, equal in length. Receptacle naked. Florets yellow. Achene ± fusiform, slightly compressed, with 5 main ribs alternating with 1 or 2 secondary ribs, usually scabrid of antrorse acute papillae especially toward apex, rarely glabrous or muriculate, apex attenuate or with a beak less than 1/5 or to 1/2 of achene length. Pappus white, scabrid, usually ± caducous.

About 15 species: C and E Asia, including N Pacific Bonin (Ogasawara) Islands; nine species (two endemic) in China.

Crepidiastrum, in the circumscription used here, includes Paraixeris, following the conclusions by Pak and Kawano (Mem. Fac. Sci. Kyoto Univ., Ser. Biol. 15: 29-61. 1992) of their carpological and cytological investigations, which have been corroborated through recent molecular phylogenetic analyses by J. W. Zhang et al. (in prep.). The analyses by J. W. Zhang et al., moreover, revealed that the Youngia segregate Crepidifolium is also nested within the Crepidiastrum clade, thus confirming an earlier assumption by Sennikov (Bot. Zhurn. 82(5): 113-116. 1997), which Sennikov later revised in favor of establishing the separate genus Crepidifolium (Sennikov & I. D. Illarionova, Komarovia 5: 96. 2008). This genus is therefore also treated here as a congener of Crepidiastrum, extending its geographical range to C Asia. Its basic chromosome number is x = 5.

1 Inner phyllaries 5; florets 5 or 6.   5 C. chelidoniifolium
+ Inner phyllaries (7 or)8; florets 8-19   (2)
2 (1) Stem leaves never clasping   (3)
+ At least upper stem leaves conspicuously clasping   (6)
3 (2) Plant stoloniferous; leaf blade triangular to orbicular.   6 C. humifusum
+ Plant not stoloniferous; leaf blade oblanceolate, elliptic, or lanceolate   (4)
4 (3) Stems numerous, intricately and divaricately dichotomously branched.   9 C. akagii
+ Stems 1 or few, paniculately branched   (5)
5 (4) Involucre 0.8-1.1 cm, 2.5-3.5 mm wide at anthesis (in middle third); anther tube 3.5-5 mm; achene black to blackish.   7 C. tenuifolium
+ Involucre 1-1.4 cm, (3.5-)4-6 mm wide at anthesis (in middle third); anther tube 5-6 mm; achene usually brownish.   8 C. diversifolium
6 (2) Annual or biennial herbs; achene with a beak 1/5-1/3 as long as achene   (7)
+ Perennial rosulate herbs with a woody caudex or subshrubs; achene without a beak or with a beak at most 1/5 as long as achene   (8)
7 (6) Middle and upper stem leaves broadest in basal third; involucre 4.5-6.5 mm; anther tube and style pure yellow upon drying.   1 C. sonchifolium
+ Middle and upper stem leaves broadest in middle third; involucre 6-9 mm; anther tube and style greenish to blackish upon drying.   4 C. denticulatum
8 (6) Leaves undivided or pinnately lobed, margin entire to shallowly dentate; primary stems decumbent; middle and upper stem leaves obovate, ovate, or lanceolate, apex subacute to rounded.   2 C. lanceolatum
+ Leaves always undivided, margin entire to denticulate; primary stems ascending-erect; upper stem leaves broadly elliptic to broadly ovate, apex mostly rounded.   3 C. taiwanianum

  • List of lower taxa


  • Flora of China @
    Browse by
    Advanced Search

    Flora of China Home




     |  eFlora Home |  People Search  |  Help  |  ActKey  |  Hu Cards  |  Glossary  |