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FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae

2. Ctenitis (C. Christensen) C. Christensen in Verdoorn, Man. Pteridol. 544. 1938.

肋毛蕨属 lei mao jue shu

Authors: Shiyong Dong & Maarten J. M. Christenhusz

Dryopteris subg. Ctenitis C. Christensen, Biol. Arbejder Tilegnede. Eug. Warming, 77. 1911; Ataxipteris Holttum.

Plants terrestrial, 10-140 cm tall. Rhizome short, erect to ascending, densely scaly at apex and at stipe bases. Fronds clustered; stipe dark brown to stramineous, scaly throughout; scales of two different types: (1) lanceolate, membranous, appressed, and iridescent scales, and (2) subulate, stiff, spreading scales without obvious iridescence. Lamina oblong-lanceolate to ovate-triangular, broadest at base, 1-4-pinnate; basal pair of pinnae always largest, triangular or broadly lanceolate, basal basiscopic pinnules of basal pinnae usually much enlarged; distal pinnae or pinnules ± adnate to rachis or costae and ± decurrent at their bases; veins all free, veinlets on ultimate pinnules or lobes simple or (rarely) bifurcate to pinnate; lamina texture herbaceous or papery, rarely leathery, with ctenitoid hairs on adaxial surface and short glandular hairs on abaxial surface, or rarely glabrous; costae of pinnules prominent, always covered with ctenitoid (multicellular) hairs. Sori medial or rarely submarginal; indusia present or not, sometimes very small and hidden by maturing sporangia. Perispore echinate, coarsely folded, or tuberculate. x = 41.

About 100-150 species: tropics and subtropics of America, Africa, Asia, and Australia; ten species (four endemic) in China.

1 Scales on stipe, rachis, or at least those on abaxial surface of costae lanceolate to ovate-lanceolate   (2)
+ Scales on stipe, rachis, and on abaxial surface of costae linear-lanceolate to subulate (inrolled and hairlike) or nearly so   (5)
2 (1) Pinnae (except basal one) 1-pinnatifid; veinlets obscure, some veinlets between pinna lobes arising from costae; perispore coarsely folded.   9 C. sinii
+ Pinnae (except basal one) 2-pinnatifid; veinlets distinct, no veinlets arising from costae; perispore coarsely folded or echinate   (3)
3 (2) Basal pinnae broadly lanceolate with symmetrical bases; perispore coarsely folded.   5 C. guidianensis
+ Basal pinnae deltoid with asymmetrical bases, basiscopic 2 or 3 pinnules much enlarged; perispore echinate   (4)
4 (3) Sori indusiate, indusia usually fugacious or not well developed; widespread in SE Asia.   10 C. subglandulosa
+ Sori exindusiate, usually covered by scales; Hunan and Sichuan (endemic).   8 C. pseudorhodolepis
5 (1) Lamina with 2 or 3 pairs of free pinnae; pinnae shallowly lobed; some veinlets between pinna lobes arising from costae; perispore coarsely folded.   2 C. dianguiensis
+ Lamina with 5-15 pairs of free pinnae; pinnae 1-3-pinnate; no veinlets or rarely a few veinlets arising from costae; perispore coarsely folded, tuberculate, or echinate   (6)
6 (5) Stipe scales 1-3 mm, linear to subulate; sori exindusiate; perispore echinate   (7)
+ Stipe scales 3-8 mm, subulate; indusia persistent or partly fugacious; perispore coarsely folded or tuberculate   (8)
7 (6) Sori usually covered by scales.   3 C. dingnanensis
+ Sori naked, not scaly.   6 C. jinfoshanensis
8 (6) Free pinnae (except basal ones) of same size, 1-pinnatifid, pinna bases not broadened.   7 C. mannii
+ Free pinnae gradually reduced toward lamina apex, 1- or 2-pinnatifid, nearly all pinnae with broadened bases   (9)
9 (8) Fronds > 1 m, 4-pinnate at base; stipe scales and rachis rufescent, 6-8 mm.   1 C. decurrentipinnata
+ Fronds < 60 cm, 2- or 3-pinnate at base; stipe scales and rachis dark brown, 3-4 mm.   4 C. eatonii

Lower Taxa


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