1 |
Leaves 2- or 3-pinnate; leaflets dichotomously forked. |
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(2) |
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Leaves 1-pinnate; leaflets simple. |
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(4) |
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2 (1) |
Leaves 2-pinnate; leaflets dichotomously 1(-3)-forked into linear, often irregular segments (ultimate leaflets), each 10-38 cm with apex attenuate to long acuminate |
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3 Cycas micholitzii |
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Leaves 3-pinnate; secondary leaflets dichotomously (2 or)3-5-forked into linear, regular segments. |
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(3) |
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3 (2) |
Leaves 1(or 2), to 7 m; segments obovate-linear, apex shortly acuminate to caudate; sterile blade of megasporophylls with 25-35 subulate lobes |
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1 Cycas multipinnata |
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Leaves (3-)5-11(-15), to 2.7 m; segments linear, apex long attenuate or long acuminate; sterile blade of megasporophylls with 39-51 filiform lobes |
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2 Cycas debaoensis |
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4 (1) |
Leaves strongly V-shaped in cross section, longitudinally recurved; margin of leaflets revolute (slightly recurved in C. taitungensis). |
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(5) |
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Leaves flat to inconspicuously V-shaped in cross section, longitudinally flat; margin of leaflets not revolute (except in C. ferruginea). |
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(7) |
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5 (4) |
Trunk not tomentose at apex; bark pale gray, smooth toward base of trunk; mature leaflets glaucous |
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15 Cycas hongheensis |
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Trunk densely tomentose at apex; bark black, scaly; mature leaflets not glaucous. |
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(6) |
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6 (5) |
Leaflets strongly recurved along margin; sterile blade of megasporophylls ovate to narrowly so, deeply laciniate; sclerotesta of seed not grooved on sides |
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6 Cycas revoluta |
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Leaflets flat or slightly recurved along margin; sterile blade of megasporophyll suborbicular or rhombic-orbicular, pectinate; sclerotesta of seed with 2 or 3 irregular grooves on each side |
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7 Cycas taitungensis |
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7 (4) |
Trunk subterranean, base not swollen; median leaflets (12-)18-25 mm wide, margin often undulate; median microsporophylls blunt. |
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(8) |
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Trunk not subterranean (or, if so, then base swollen); median leaflets 6-11(-14) mm wide, margin flat (revolute in C. ferruginea); median microsporophylls spinose. |
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(9) |
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8 (7) |
Petioles green in 1st year; megasporophylls 5-15(-20), loosely grouped, sterile blade divided into 15-25 lobes |
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4 Cycas balansae |
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Petioles blue in 1st year, later green; megasporophylls 25-50, densely grouped, divided into 33-45 lobes |
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5 Cycas segmentifida |
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9 (7) |
Leaves (0.7-)1-3 m × 30-60 cm; terminal lobe of sterile blade of megasporophyll usually flattened. |
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(10) |
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Leaves 0.5-1(-1.3) m × (12-)20-30(-40) cm; terminal lobe of sterile blade of megasporophyll subulate, not flattened. |
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(12) |
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10 (9) |
Trunk inflated; bark white-gray, nearly smooth toward base of trunk; median leaflets often less than 10 mm wide; petiole densely spinose from base upward; seeds 3.5-4.5 cm |
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9 Cycas hainanensis |
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Trunk cylindric; bark dark, scaly; median leaflets often more than 11 mm wide; petiole without spines or with sparse spines near base; seeds 2.5-3 cm. |
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(11) |
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11 (10) |
Leaflets longitudinally inserted at 55-75° to rachis; apex of cataphylls with hard point; sterile blade of megasporophyll rhombic-ovate; sclerotesta of seed finely verrucose |
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10 Cycas taiwaniana |
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Leaflets longitudinally inserted at ca. 50° to rachis; apex of cataphylls with soft point; sterile blade of megasporophyll broadly ovate, obovate, or suborbicular; sclerotesta of seed smooth |
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11 Cycas szechuanensis |
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12 (9) |
Trunk cylindric, 1-7 m; bark white-gray, gray, or brown, scaly or raised; pollen cones (25-)30-45 × 8-15 cm, microsporophylls 3.5-6 cm; megasporophylls 13-20 cm. |
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(13) |
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Trunk subterranean, base swollen; bark gray, smooth; pollen cones 20-30 × 6-8 cm, microsporophylls 1.5-3 cm; megasporophylls 8-14 cm. |
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(14) |
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13 (12) |
Trunk often dichotomously branched toward apex, to 16 m, apex not tomentose; bark gray or white-gray, smooth toward base of trunk; leaflets with midvein sulcate adaxially and margin revolute to slightly recurved; seeds 4.5-6 × 4-4.7 cm |
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16 Cycas pectinata |
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Trunk simple, to 2(-3) m, apex densely tomentose; bark brown to dark gray, scaly; leaflets with midvein never sulcate adaxially and margin flat to slightly recurved; seeds 2.5-3.5 × 2.2-3 cm |
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8 Cycas panzhihuaensis |
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14 (12) |
Leaflets longitudinally inserted at 55-65° to rachis; petiole with 9-16 spines along each side throughout length (SW Hainan) |
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14 Cycas changjiangensis |
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Leaflets longitudinally inserted at ca. 90° to rachis; petiole with 0-8(or more) spines along each side often only in apical part (W Guangxi). |
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(15) |
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15 (14) |
Young leaves densely rusty brown tomentose abaxially; leaflets not overlapping, light green at maturity, 20-30 cm × 5-12 mm, base attenuate, margin strongly revolute |
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12 Cycas ferruginea |
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Young leaves sparsely red-brown tomentose abaxially; leaflets often overlapping, dark green at maturity, 10-18 cm × 14-18 mm, base usually truncate, margin flat or only slightly revolute |
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13 Cycas miquelii |
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