1 |
Leaf blade pinnately compound; plants annual; petals enlarged distally; seeds transversely lamellate (5. D. sect. Anthriscifolium) |
|
173 D. anthriscifolium |
+ |
Leaf blade palmately parted (simple or compound); plants perennial; petals not enlarged distally; seeds narrowly winged along angles, or transversely rugose or squamulose winged. |
|
(2) |
|
|
|
|
2 (1) |
Sepals and staminodes yellow (4. D. sect. Oligophyllon) |
|
172 D. biternatum |
+ |
Sepals and staminodes blue, purple, black, brown, or white (if sepals yellow, then staminodes black). |
|
(3) |
|
|
|
|
3 (2) |
Petals and staminodes wholly or partly black to brown. |
|
(4) |
+ |
Petals and staminodes not black to brown, similar in color to sepals (blue, purple, or white) (3. D. sect. Delphinastrum). |
|
(78) |
|
|
|
|
4 (3) |
Staminode limb weakly differentiated from claw (limb less than 2 × as wide as claw and less than 45° angle from claw), adaxially glabrous or ciliate (1. D. sect. Aconitoides). |
|
(5) |
+ |
Staminode limb distinct from claw in width (limb at least 2 × as wide as claw) and orientation (limb at least 45° from claw), usually adaxially barbate (2. D. sect. Elatopsis). |
|
(10) |
|
|
|
|
5 (4) |
Spur pendulous; yellow, glandular hairs absent from stem. |
|
(6) |
+ |
Spur within 45° of horizontal; yellow, glandular hairs present on stem. |
|
(8) |
|
|
|
|
6 (5) |
Inflorescence tapered cylindric, usually at least 16 cm |
|
6 D. trichophorum |
+ |
Inflorescence ovoid to globose, usually less than 16 cm. |
|
(7) |
|
|
|
|
7 (6) |
Spur less than 1.7 cm |
|
1 D. forrestii |
+ |
Spur at least 1.7 cm |
|
5 D. oxycentrum |
|
|
|
|
8 (5) |
Stem less than 20 cm; inflorescence less than 3 × as long as wide, ca. 4-flowered |
|
3 D. xichangense |
+ |
Stem 20--50(--70) cm; inflorescence at least 3 × as long as wide, (6--)10--20-flowered. |
|
(9) |
|
|
|
|
9 (8) |
Bracteoles at least 1.2 cm; raceme dense |
|
2 D. leiophyllum |
+ |
Bracteoles less than 1.2 cm; raceme lax |
|
4 D. wrightii |
|
|
|
|
10 (4) |
Inflorescence less than 3 × as long as wide, usually 1- or few flowered (to 40-flowered in D. densiflorum). |
|
(11) |
+ |
Inflorescence at least 3 × as long as wide, usually more than 6-flowered. |
|
(47) |
|
|
|
|
11 (10) |
Carpels 4 or 5. |
|
(12) |
+ |
Carpels 3. |
|
(20) |
|
|
|
|
12 (11) |
Leaf base abruptly narrowed and cuneate |
|
14 D. brunonianum |
+ |
Leaf base not narrowed. |
|
(13) |
|
|
|
|
13 (12) |
Leaf lobes separate for less than 90% of blade radius. |
|
(14) |
+ |
Leaf lobes separate for at least 90% of blade radius. |
|
(16) |
|
|
|
|
14 (13) |
Spur subulate, ca. 2.5 mm in diam. at base |
|
34 D. lingbaoense |
+ |
Spur conic, (3--)4--7 mm in diam. at base. |
|
(15) |
|
|
|
|
15 (14) |
Stem 6--10 cm; spur less than 9 mm |
|
13 D. lacostei |
+ |
Stem 40--50 cm; spur more than 9 mm |
|
15 D. yechengense |
|
|
|
|
16 (13) |
Flower solitary; carpels 4 |
|
20 D. chumulangmaense |
+ |
Flowers 2--6; carpels 5 (rarely 4). |
|
(17) |
|
|
|
|
17 (16) |
Spur conic or narrowly so, less than 2 cm, base 4--5 mm in diam. |
|
(18) |
+ |
Spur subulate or cylindric-subulate, more than 2 cm, base ca. 3 mm in diam. |
|
(19) |
|
|
|
|
18 (17) |
Filaments glabrous |
|
17 D. glaciale |
+ |
Filaments pilose |
|
18 D. pseudoglaciale |
|
|
|
|
19 (17) |
Inflorescence with yellow, glandular hairs; petal apex emarginate; staminode limb suboblong, 2-cleft; filaments glabrous |
|
32 D. pylzowii |
+ |
Inflorescence without glandular hairs; petal apex entire; staminode limb suborbicular, emarginate; filaments sparsely pilose |
|
33 D. motingshanicum |
|
|
|
|
20 (11) |
Leaf lobes separate for less than 90% of blade radius. |
|
(21) |
+ |
Leaf lobes separate for at least 90% of blade radius. |
|
(33) |
|
|
|
|
21 (20) |
Carpels hairy only on ventral suture, glabrous elsewhere |
|
24 D. wardii |
+ |
Carpels evenly hairy throughout exposed surfaces. |
|
(22) |
|
|
|
|
22 (21) |
Spur conic or conic-tubular, base 5--12 mm in diam. |
|
(23) |
+ |
Spur cylindric or subulate, base 2--4(--4.5) mm in diam. |
|
(25) |
|
|
|
|
23 (22) |
Bracteoles blue-purple, navicular-lanceolate |
|
41 D. purpurascens |
+ |
Bracteoles not blue-purple, flat. |
|
(24) |
|
|
|
|
24 (23) |
Bracteoles more than 0.5 mm from flowers; spur base 7--12 mm in diam. |
|
16 D. chrysotrichum |
+ |
Bracteoles ca. 0.5 mm from flowers; spur base ca. 5 mm in diam. |
|
19 D. viscosum |
|
|
|
|
25 (22) |
Bracteoles ca. 1 mm from flowers. |
|
(26) |
+ |
Bracteoles more than 1 mm from flowers. |
|
(29) |
|
|
|
|
26 (25) |
Bracteoles elliptic |
|
39 D. albocoeruleum |
+ |
Bracteoles linear or narrowly so. |
|
(27) |
|
|
|
|
27 (26) |
Staminodes yellow barbate |
|
27 D. chenii |
+ |
Staminodes white barbate. |
|
(28) |
|
|
|
|
28 (27) |
Spur less than 1 cm |
|
12 D. pulanense |
+ |
Spur more than 1 cm |
|
61 D. wentsaii |
|
|
|
|
29 (25) |
Staminode limb emarginate at apex |
|
28 D. yangii |
+ |
Staminode limb 2-lobed to middle. |
|
(30) |
|
|
|
|
30 (29) |
Sepal margin white |
|
27 D. chenii |
+ |
Sepal margin not white. |
|
(31) |
|
|
|
|
31 (30) |
Lateral and lower sepals obovate or broadly so |
|
30 D. pseudopulcherrimum |
+ |
Lateral and lower sepals elliptic or ovate. |
|
(32) |
|
|
|
|
32 (31) |
Stem less than 50 cm |
|
25 D. smithianum |
+ |
Stem ca. 50 cm |
|
40 D. siwanense |
|
|
|
|
33 (20) |
Bracteoles ca. 1 mm from flower. |
|
(34) |
+ |
Bracteoles more than 1 mm from flower. |
|
(35) |
|
|
|
|
34 (33) |
Bracteoles linear, 1--1.7 mm wide |
|
29 D. batangense |
+ |
Bracteoles narrowly elliptic, 3--3.5 mm wide |
|
38 D. henryi |
|
|
|
|
35 (33) |
Bracteoles leaflike, 3-cleft. |
|
(36) |
+ |
Bracteoles undivided, linear or lanceolate. |
|
(40) |
|
|
|
|
36 (35) |
Spur longer than lateral sepals. |
|
(37) |
+ |
Spur shorter than or equaling lateral sepals. |
|
(38) |
|
|
|
|
37 (36) |
Sepals 2--3 cm, spur subulate |
|
35 D. candelabrum |
+ |
Sepals 1.4--1.5 cm, spur cylindric |
|
37 D. calophyllum |
|
|
|
|
38 (36) |
Inflorescence 1-flowered; staminodes undivided |
|
23 D. chungbaense |
+ |
Inflorescence 2- or 3-flowered; staminodes 2-lobed. |
|
(39) |
|
|
|
|
39 (38) |
Sepals 1.9--2 cm, spur subcylindric, shorter than sepals, ca. 1.2 cm; filaments pilose |
|
36 D. pseudocandelabrum |
+ |
Sepals 1.2--1.5 cm, spur subulate, subequaling sepals, 1.2--1.4 cm; filaments glabrous |
|
22 D. qinghaiense |
|
|
|
|
40 (35) |
Inflorescence 1-flowered; spur shorter than lateral sepals. |
|
(41) |
+ |
Inflorescence 2--7-flowered; spur longer than lateral sepals. |
|
(42) |
|
|
|
|
41 (40) |
Spur cylindric, 1--1.3 cm |
|
21 D. tangkulaense |
+ |
Spur cylindric-subulate or subulate, 1.7--2.6 cm |
|
35 D. candelabrum |
|
|
|
|
42 (40) |
Lower sepals obovate, apex rounded |
|
30 D. pseudopulcherrimum |
+ |
Lower sepals ovate, elliptic-ovate, or narrowly elliptic, apex narrowed. |
|
(43) |
|
|
|
|
43 (42) |
Central leaf segment 2- or 3-subpinnately dissected, ultimate lobules narrowly ovate to linear |
|
32 D. pylzowii |
+ |
Central leaf segment 3-parted, ultimate lobules ovate or narrowly so. |
|
(44) |
|
|
|
|
44 (43) |
Lateral sepals equaling or longer than spur; seeds not densely squamulose winged |
|
22 D. qinghaiense |
+ |
Lateral sepals shorter than spur or, if equaling spur (D. siwanense), seeds densely squamulose winged. |
|
(45) |
|
|
|
|
45 (44) |
Sepals 2--2.2 cm, spur 3.5--3.8 cm |
|
26 D. malacophyllum |
+ |
Sepals 1.1--1.8 cm, spur 1.6--2.4 cm. |
|
(46) |
|
|
|
|
46 (45) |
Stem 28--30 cm; sepal margin white; spur 2--2.4 cm |
|
31 D. yajiangense |
+ |
Stem 38--120 cm; sepal margin not white; spur 1.6--1.8 cm |
|
40 D. siwanense |
|
|
|
|
47 (10) |
Spur ca. 10 mm, much shorter than lateral sepals. |
|
(48) |
+ |
Spur at least 9 mm, equaling or longer than lateral sepals, rarely slightly shorter than them (D. subspathulatum, D. vestitum, and D. wuqiaense). |
|
(52) |
|
|
|
|
48 (47) |
Carpels pubescent. |
|
(49) |
+ |
Carpels glabrous. |
|
(51) |
|
|
|
|
49 (48) |
Basal leaves absent at anthesis |
|
8 D. nordhagenii |
+ |
Basal leaves present at anthesis. |
|
(50) |
|
|
|
|
50 (49) |
Raceme 20--40-flowered |
|
7 D. densiflorum |
+ |
Raceme less than 20-flowered |
|
12 D. pulanense |
|
|
|
|
51 (48) |
Stem 30--38 cm; bracteoles sessile, linear-lanceolate |
|
9 D. tetragynum |
+ |
Stem 20--23 cm; bracteoles shortly stalked, obovate, elliptic, or linear |
|
10 D. taxkorganense |
|
|
|
|
52 (47) |
Leaves all basal, or 1 or 2 proximal cauline leaves present. |
|
(53) |
+ |
Leaves both basal and cauline, cauline leaves evenly distributed. |
|
(57) |
|
|
|
|
53 (52) |
Leaf blade cleft to base, ultimate lobules linear to narrowly lanceolate |
|
67 D. erlangshanicum |
+ |
Leaf blade cleft less than 80% to base, ultimate lobules not linear. |
|
(54) |
|
|
|
|
54 (53) |
Hairs of bracteole margin 1.5--2.2 mm |
|
46 D. iliense |
+ |
Hairs of bracteole margin ca. 0.8 mm. |
|
(55) |
|
|
|
|
55 (54) |
Rachis with yellow, glandular hairs |
|
57 D. kaschgaricum |
+ |
Rachis without yellow, glandular hairs. |
|
(56) |
|
|
|
|
56 (55) |
Lateral sepals at least 1.2 cm |
|
58 D. zhangii |
+ |
Lateral sepals less than 1.2 cm |
|
44 D. kansuense |
|
|
|
|
57 (52) |
Leaf blade lobed less than 50% to base. |
|
(58) |
+ |
Leaf blade lobed more than 50% to base. |
|
(59) |
|
|
|
|
58 (57) |
Stem without glandular hairs; spur cylindric-conic, slightly shorter than lateral sepals, 1.1--1.3 cm, base 5--6 mm in diam. |
|
11 D. vestitum |
+ |
Stem with yellow, glandular hairs; spur subulate, slightly longer than lateral sepals, 1.9--2.6 cm, base 3--4 mm in diam. |
|
43 D. nangziense |
|
|
|
|
59 (57) |
Bracteoles borne proximally on pedicel |
|
51 D. maackianum |
+ |
Bracteoles borne near middle of pedicel or distally. |
|
(60) |
|
|
|
|
60 (59) |
Bracteoles navicular-ovate |
|
48 D. naviculare |
+ |
Bracteoles flat, not navicular. |
|
(61) |
|
|
|
|
61 (60) |
Pedicels glabrous. |
|
(62) |
+ |
Pedicels hairy. |
|
(65) |
|
|
|
|
62 (61) |
Sepals abaxially densely puberulent |
|
62 D. mollifolium |
+ |
Sepals abaxially glabrous. |
|
(63) |
|
|
|
|
63 (62) |
Bracteoles contiguous to flowers |
|
65 D. korshinskyanum |
+ |
Bracteoles remote from flowers. |
|
(64) |
|
|
|
|
64 (63) |
Bracteoles narrowly lanceolate or linear-oblanceolate, margin long ciliate |
|
46 D. iliense |
+ |
Bracteoles narrowly linear, margin not ciliate |
|
59 D. sinoelatum |
|
|
|
|
65 (61) |
Bracteoles 2--3 mm wide. |
|
(66) |
+ |
Bracteoles ca. 1 mm wide. |
|
(70) |
|
|
|
|
66 (65) |
Stem with dense, yellow, glandular hairs |
|
45 D. hillcoatiae |
+ |
Stem without yellow, glandular hairs. |
|
(67) |
|
|
|
|
67 (66) |
Bracteoles remote from flowers. |
|
(68) |
+ |
Bracteoles contiguous to flowers. |
|
(69) |
|
|
|
|
68 (67) |
Stem simple; pedicels to 8 cm; spur slightly shorter than sepals |
|
42 D. subspathulatum |
+ |
Stem branched; pedicels to 2.7 cm; spur slightly longer than sepals |
|
49 D. ellipticovatum |
|
|
|
|
69 (67) |
Raceme rachis without glandular hairs; bracteoles lanceolate-oblong or lanceolate, 8--10 mm |
|
50 D. winklerianum |
+ |
Raceme rachis with yellow, glandular hairs; bracteoles linear, 10--14 mm |
|
64 D. tabatae |
|
|
|
|
70 (65) |
Sepals abaxially without yellow, glandular hairs. |
|
(71) |
+ |
Sepals abaxially with yellow, glandular hairs. |
|
(74) |
|
|
|
|
71 (70) |
Carpels glabrous |
|
63 D. eglandulosum |
+ |
Carpels pubescent. |
|
(72) |
|
|
|
|
72 (71) |
Pedicels and sepals abaxially strigose; carpels densely strigose |
|
54 D. tianshanicum |
+ |
Pedicels and sepals abaxially puberulent; carpels puberulent. |
|
(73) |
|
|
|
|
73 (72) |
Petals sparsely ciliate, apex 2-lobed |
|
47 D. aktoense |
+ |
Petals glabrous, apex emarginate |
|
66 D. trisectum |
|
|
|
|
74 (70) |
Central primary leaf lobes and secondary lobes of lateral primary lobes long acuminate or caudate at apex |
|
60 D. shawurense |
+ |
Central primary leaf lobes and secondary lobes of lateral primary lobes acute or acuminate at apex. |
|
(75) |
|
|
|
|
75 (74) |
Bracteoles contiguous to flowers. |
|
(76) |
+ |
Bracteoles remote from flowers. |
|
(77) |
|
|
|
|
76 (75) |
Petals 2-lobed; carpels densely puberulent |
|
52 D. tarbagataicum |
+ |
Petals entire; carpels glabrous |
|
53 D. elatum |
|
|
|
|
77 (75) |
Spur shorter than or equaling lateral sepals |
|
55 D. wuqiaense |
+ |
Spur longer than lateral sepals |
|
56 D. kunlunshanicum |
|
|
|
|
78 (3) |
Cauline leaf petiole sheath 8--15 mm wide |
|
139 D. coleopodum |
+ |
Cauline leaf petiole sheath less than 8 mm wide, or cauline leaves absent. |
|
(79) |
|
|
|
|
79 (78) |
Carpels at least 4. |
|
(80) |
+ |
Carpels 3. |
|
(88) |
|
|
|
|
80 (79) |
Staminode limb cleft for at least 30% of its length. |
|
(81) |
+ |
Staminode limb cleft for less than 30% of its length. |
|
(84) |
|
|
|
|
81 (80) |
Inflorescence less than 3 × as long as wide |
|
162 D. caeruleum |
+ |
Inflorescence at least 3 × as long as wide. |
|
(82) |
|
|
|
|
82 (81) |
Stem at least 40 cm |
|
154 D. pseudomosoynense |
+ |
Stem less than 40 cm. |
|
(83) |
|
|
|
|
83 (82) |
Spur less than 1 cm |
|
8 D. nordhagenii |
+ |
Spur at least 1 cm |
|
69 D. humilius |
|
|
|
|
84 (80) |
Inflorescence more than 3 × as long as wide. |
|
(85) |
+ |
Inflorescence less than 3 × as long as wide. |
|
(86) |
|
|
|
|
85 (84) |
Staminode limb white barbate |
|
68 D. kingianum |
+ |
Staminode limb yellow barbate |
|
153 D. sinopentagynum |
|
|
|
|
86 (84) |
Staminode limb narrowly obovate; spur at least 2.4 cm, base at least 3 mm in diam. |
|
161 D. beesianum |
+ |
Staminode limb broadly obovate or suborbicular; spur to 2.6 cm, base to 4 mm in diam. |
|
(87) |
|
|
|
|
87 (86) |
Staminodes white barbate |
|
163 D. huangzhongense |
+ |
Staminodes yellow barbate |
|
162 D. caeruleum |
|
|
|
|
88 (79) |
Sepals abaxially corniculate near apex. |
|
(89) |
+ |
Sepals abaxially not or very shortly (less than 0.5 mm) corniculate near apex. |
|
(97) |
|
|
|
|
89 (88) |
Corniculate projection 2.5--3.5 mm, 2-cleft |
|
105 D. campylocentrum |
+ |
Corniculate projection less than 2.5 mm, not 2-cleft. |
|
(90) |
|
|
|
|
90 (89) |
Spur ca. 1.3 cm |
|
111 D. sparsiflorum |
+ |
Spur more than 1.3 cm. |
|
(91) |
|
|
|
|
91 (90) |
Sepals yellow pubescent abaxially |
|
132 D. leiostachyum |
+ |
Sepals white pubescent abaxially. |
|
(92) |
|
|
|
|
92 (91) |
Inflorescence pubescent |
|
106 D. sutchuenense |
+ |
Inflorescence glabrous. |
|
(93) |
|
|
|
|
93 (92) |
Petals apically 2-lobate |
|
117 D. taliense |
+ |
Petals apically entire or emarginate. |
|
(94) |
|
|
|
|
94 (93) |
Spur ca. 2.4 cm. |
|
(95) |
+ |
Spur less than 2.4 cm. |
|
(96) |
|
|
|
|
95 (94) |
Filaments pilose |
|
120 D. ceratophoroides |
+ |
Filaments glabrous |
|
131 D. dolichocentroides |
|
|
|
|
96 (94) |
Basal leaves present at anthesis |
|
128 D. ceratophorum |
+ |
Basal leaves absent at anthesis |
|
136 D. yunnanense |
|
|
|
|
97 (88) |
Spur U-shaped or spirally recurved (more than 180°). |
|
(98) |
+ |
Spur straight, upcurved, or recurved to 180°, neither U-shaped nor spirally curved. |
|
(107) |
|
|
|
|
98 (97) |
Pedicels glabrous. |
|
(99) |
+ |
Pedicels hairy. |
|
(101) |
|
|
|
|
99 (98) |
Inflorescence less than 3 × as long as wide; leaf blade lobed ca. 50% to base |
|
96 D. maoxianense |
+ |
Inflorescence at least 3 × as long as wide; leaf blade lobed more than 50% to base. |
|
(100) |
|
|
|
|
100 (99) |
Leaf blade lobed less than 90% to base |
|
92 D. potaninii |
+ |
Leaf blade lobed at least 90% to base |
|
113 D. glabricaule |
|
|
|
|
101 (98) |
Leaf blade lobed ca. 50% to base |
|
95 D. sinovitifolium |
+ |
Leaf blade lobed more than 50% to base. |
|
(102) |
|
|
|
|
102 (101) |
Leaf lobes long acuminate or caudate at apex |
|
103 D. pseudotongolense |
+ |
Leaf lobes shortly acuminate at apex. |
|
(103) |
|
|
|
|
103 (102) |
Stem proximally glabrous. |
|
(104) |
+ |
Stem proximally hairy. |
|
(105) |
|
|
|
|
104 (103) |
Raceme with only yellow, glandular hairs; petals undivided |
|
100 D. wenchuanense |
+ |
Raceme with both appressed, white hairs and yellow, glandular hairs; petals 2-lobed |
|
101 D. bulleyanum |
|
|
|
|
105 (103) |
Stem sparsely pubescent proximally |
|
99 D. angustirhombicum |
+ |
Stem hispid proximally. |
|
(106) |
|
|
|
|
106 (105) |
Leaf blade strongly divided, with many lobules; lateral sepals 1.5--1.8 cm; carpels hairy |
|
97 D. spirocentrum |
+ |
Leaf blade less divided, with few lobules; lateral sepals ca. 1.1 cm; carpels glabrous |
|
98 D. gonggaense |
|
|
|
|
107 (97) |
Leaf blade lobed up to 90% to base. |
|
(108) |
+ |
Leaf blade lobed more than 90% to base. |
|
(164) |
|
|
|
|
108 (107) |
Sepals hairy on both surfaces. |
|
(109) |
+ |
Sepals adaxially glabrous. |
|
(110) |
|
|
|
|
109 (108) |
Stem, raceme rachis, and pedicels densely appressed puberulent; bracteoles 10--18 mm; spur subcylindric |
|
70 D. pachycentrum |
+ |
Stem and raceme rachis glabrous, pedicels glabrous proximally, puberulent distally; bracteoles ca. 5 mm; spur subulate |
|
75 D. yuchuanii |
|
|
|
|
110 (108) |
Spur shorter than or equaling lateral sepals. |
|
(111) |
+ |
Spur longer than lateral sepals, rarely subequaling them. |
|
(116) |
|
|
|
|
111 (110) |
Inflorescence less than 3 × as long as wide |
|
92 D. potaninii |
+ |
Inflorescence ca. 3 × as long as wide. |
|
(112) |
|
|
|
|
112 (111) |
Spur ca. 9 mm, upcurved |
|
77 D. orthocentrum |
+ |
Spur more than 10 mm, usually horizontal or recurved. |
|
(113) |
|
|
|
|
113 (112) |
Leaf primary lobes cleft ca. 50% of blade radius |
|
116 D. aemulans |
+ |
Leaf primary lobes cleft less than 50% of blade radius. |
|
(114) |
|
|
|
|
114 (113) |
Bracteoles less than 0.8 mm wide |
|
88 D. longipedicellatum |
+ |
Bracteoles 1.2--2 mm wide. |
|
(115) |
|
|
|
|
115 (114) |
Bracteoles more than 0.5 mm from flowers |
|
72 D. gyalanum |
+ |
Bracteoles ca. 0.5 mm from flowers |
|
76 D. micropetalum |
|
|
|
|
116 (110) |
Bracteoles ca. 1 mm from flowers. |
|
(117) |
+ |
Bracteoles more than 1 mm from flowers. |
|
(121) |
|
|
|
|
117 (116) |
Basal leaves 4--8, present at anthesis; cauline leaves 1--3, borne proximally on stem. |
|
(118) |
+ |
Basal leaves usually wilted at anthesis; cauline leaves at least 3, evenly arranged along stem. |
|
(119) |
|
|
|
|
118 (117) |
Stem ca. 40 cm, spreading hispid throughout; raceme ca. 20 cm; spur 1.7--2 cm |
|
127 D. yongningense |
+ |
Stem less than 40 cm, sparsely strigose near base, glabrous elsewhere; raceme less than 20 cm; spur 2.4--2.6 cm |
|
133 D. brevisepalum |
|
|
|
|
119 (117) |
Bracteoles narrowly lanceolate |
|
79 D. delavayi |
+ |
Bracteoles linear. |
|
(120) |
|
|
|
|
120 (119) |
Spur more than 1.8 cm; pedicels more than 1 cm |
|
80 D. umbrosum |
+ |
Spur ca. 1.8 cm; pedicels ca. 1 cm |
|
89 D. pseudocyananthum |
|
|
|
|
121 (116) |
Leaf cleft less than 60% of blade radius. |
|
(122) |
+ |
Leaf cleft ca. 60% of blade radius. |
|
(124) |
|
|
|
|
122 (121) |
Raceme glabrous |
|
121 D. hamatum |
+ |
Raceme puberulent. |
|
(123) |
|
|
|
|
123 (122) |
Inflorescence yellow glandular puberulent; ovaries pubescent |
|
122 D. hueizeense |
+ |
Inflorescence white puberulent; ovaries glabrous |
|
123 D. calthifolium |
|
|
|
|
124 (121) |
Leaf blade lobed more than 85% of blade radius. |
|
(125) |
+ |
Leaf blade lobed ca. 85% of blade radius. |
|
(143) |
|
|
|
|
125 (124) |
Pedicels glabrous. |
|
(126) |
+ |
Pedicels hairy. |
|
(131) |
|
|
|
|
126 (125) |
Staminode limb 2-cleft to middle. |
|
(127) |
+ |
Staminode limb entire or emarginate at apex. |
|
(129) |
|
|
|
|
127 (126) |
Middle cauline leaves with central lobe broadly rhombic, 3-cleft; pedicels ascending-spreading, forming an obtuse angle with rachis |
|
134 D. tenii |
+ |
Middle cauline leaves with central lobe lanceolate, often undivided, or with 1 or 2 lobules; pedicels erect-spreading, forming an acute angle with rachis. |
|
(128) |
|
|
|
|
128 (127) |
Raceme laxly 3--10-flowered; spur 1.7--2.4 cm; petals glabrous, emarginate at apex |
|
136 D. yunnanense |
+ |
Raceme densely 18--30-flowered; spur 1.3--1.5 cm; petals ciliate at margin, 2-lobed at apex |
|
137 D. pseudoyunnanense |
|
|
|
|
129 (126) |
Stem at least 80 cm, ca. 10-leaved |
|
74 D. cheilanthum |
+ |
Stem less than 80 cm, up to 6-leaved. |
|
(130) |
|
|
|
|
130 (129) |
Sepals ca. 1.8 cm, spur 2.6--3 cm; staminode limb entire at apex |
|
119 D. liangshanense |
+ |
Sepals 1--1.4 cm, spur 1.8--2.5 cm; staminode limb emarginate at apex |
|
134 D. tenii |
|
|
|
|
131 (125) |
Raceme rachis glabrous. |
|
(132) |
+ |
Raceme rachis hairy. |
|
(134) |
|
|
|
|
132 (131) |
Stem ca. 70 cm; raceme to 22 cm, ca. 8-flowered |
|
156 D. obcordatilimbum |
+ |
Stem 11--30 cm; raceme 3--16 cm, 1--5-flowered. |
|
(133) |
|
|
|
|
133 (132) |
Stem ca. 30 cm; raceme 3--5-flowered; sepals 0.75--0.9 cm, spur ca. 1.9 cm; staminode limb 2-parted for more than 50% of its length |
|
85 D. yanwaense |
+ |
Stem 11--18 cm; raceme 1- or 2-flowered; sepals 1.6--1.7 cm, spur ca. 2.8 cm; staminode limb entire |
|
124 D. pseudohamatum |
|
|
|
|
134 (131) |
Pedicels without glandular hairs. |
|
(135) |
+ |
Pedicels with yellow, glandular hairs. |
|
(139) |
|
|
|
|
135 (134) |
Stem 75--95 cm; basal and lower cauline leaves wilted at anthesis. |
|
(136) |
+ |
Stem 3--17 cm; basal leaves present at anthesis. |
|
(138) |
|
|
|
|
136 (135) |
Staminode limb emarginate |
|
73 D. hsinganense |
+ |
Staminode limb 2-lobed. |
|
(137) |
|
|
|
|
137 (136) |
Bracteoles narrowly lanceolate |
|
79 D. delavayi |
+ |
Bracteoles subulate |
|
136 D. yunnanense |
|
|
|
|
138 (135) |
Stem 15--17 cm; leaf blade central lobe 3-parted; raceme 2--5-flowered |
|
129 D. yulungshanicum |
+ |
Stem 3--8 cm; leaf blade central lobe 3-dentate or 3-lobed; raceme 1- or 2-flowered |
|
130 D. hui |
|
|
|
|
139 (134) |
Leaf blade central lobe lanceolate, undivided or indistinctly 3-lobed |
|
87 D. hirtifolium |
+ |
Leaf blade central lobe rhombic or obovate-rhombic, usually distinctly 3-lobed. |
|
(140) |
|
|
|
|
140 (139) |
Leaf blade primary lobes dissected, ultimate lobules narrowly ovate to linear-lanceolate; carpels glabrous |
|
140 D. kantzeense |
+ |
Leaf blade primary lobes less divided, ultimate lobules triangular; carpels puberulent. |
|
(141) |
|
|
|
|
141 (140) |
Stem 11--30 cm; raceme 3- or 4-flowered |
|
126 D. pergameneum |
+ |
Stem more than 40 cm; raceme 5--15-flowered. |
|
(142) |
|
|
|
|
142 (141) |
Stem long hispid proximally |
|
79 D. delavayi |
+ |
Stem puberulent proximally |
|
144 D. incisolobulatum |
|
|
|
|
143 (124) |
Stem glabrous below inflorescence. |
|
(144) |
+ |
Stem hairy below inflorescence. |
|
(152) |
|
|
|
|
144 (143) |
Pedicels glabrous |
|
92 D. potaninii |
+ |
Pedicels hairy. |
|
(145) |
|
|
|
|
145 (144) |
Pedicels with only yellow, glandular hairs. |
|
(146) |
+ |
Pedicels with only white hairs or with both white hairs and yellow, glandular hairs. |
|
(147) |
|
|
|
|
146 (145) |
Inflorescence less than 3 × as long as wide |
|
93 D. eriostylum |
+ |
Inflorescence ca. 3 × as long as wide |
|
118 D. lasiantherum |
|
|
|
|
147 (145) |
Pedicels with only appressed, white hairs |
|
78 D. altissimum |
+ |
Pedicels with both appressed, white hairs and spreading, yellow, glandular hairs. |
|
(148) |
|
|
|
|
148 (147) |
Leaf blade central lobe with lateral lobules less than 2 × as long as neighboring lobules. |
|
(149) |
+ |
Leaf blade central lobe with lateral lobules ca. 2 × as long as neighboring lobules. |
|
(150) |
|
|
|
|
149 (148) |
Staminode hairs white; spur ca. 1.5 cm |
|
82 D. handelianum |
+ |
Staminode hairs yellow; spur 2--2.1 cm |
|
83 D. honanense |
|
|
|
|
150 (148) |
Leaf blade to 6.8 cm wide; raceme ca. 4-flowered; staminode limb emarginate at apex |
|
102 D. ninglangshanicum |
+ |
Leaf blade to 12 cm wide; raceme 6--16(--25)-flowered; staminode limb 2-cleft to middle. |
|
(151) |
|
|
|
|
151 (150) |
Pedicels with dense, spreading, yellow, glandular hairs and a few retrorse, white hairs; sepals 1.4--1.7 cm |
|
86 D. tongolense |
+ |
Pedicels with dense, retrorse, white hairs and sparse, spreading, yellow, glandular hairs; sepals 1--1.5 cm |
|
101 D. bulleyanum |
|
|
|
|
152 (143) |
Inflorescence less than 3 × as long as wide |
|
94 D. laxicymosum |
+ |
Inflorescence ca. 3 × as long as wide. |
|
(153) |
|
|
|
|
153 (152) |
Pedicels glabrous |
|
91 D. hirticaule |
+ |
Pedicels hairy. |
|
(154) |
|
|
|
|
154 (153) |
Leaf blade central lobes undivided or indistinctly 3-lobed. |
|
(155) |
+ |
Leaf blade central lobes distinctly 3-lobed to 3-parted. |
|
(156) |
|
|
|
|
155 (154) |
Bracteoles less than 2 mm from flowers |
|
79 D. delavayi |
+ |
Bracteoles ca. 2 mm from flowers |
|
82 D. handelianum |
|
|
|
|
156 (154) |
Stem with appressed hairs. |
|
(157) |
+ |
Stem with spreading or retrorse hairs. |
|
(158) |
|
|
|
|
157 (156) |
Stem ca. 30 cm; raceme rachis glabrous; pedicels 3.5--10 cm |
|
85 D. yanwaense |
+ |
Stem 57--65 cm; raceme rachis puberulent; pedicels 1--2.9 cm |
|
84 D. omeiense |
|
|
|
|
158 (156) |
Stem hairs 1.2--3 mm. |
|
(159) |
+ |
Stem hairs less than 1 mm or absent. |
|
(160) |
|
|
|
|
159 (158) |
Stem hairs yellowish |
|
84 D. omeiense |
+ |
Stem hairs white |
|
91 D. hirticaule |
|
|
|
|
160 (158) |
Bracteoles subulate or filiform, 2--5 × 0.2--0.4 mm, glabrous |
|
90 D. sherriffii |
+ |
Bracteoles lanceolate, 4.5--11 × 1.2--1.7 mm, hairy. |
|
(161) |
|
|
|
|
161 (160) |
Cauline leaf 1, leaf blade to 10 cm wide |
|
144 D. incisolobulatum |
+ |
Cauline leaves at least 2, leaf blade 8--16(--22) cm wide. |
|
(162) |
|
|
|
|
162 (161) |
Stem glabrous proximally; sepals less than 1 cm |
|
81 D. latirhombicum |
+ |
Stem puberulent proximally; sepals ca. 1 cm. |
|
(163) |
|
|
|
|
163 (162) |
Filaments glabrous to sparsely hairy |
|
79 D. delavayi |
+ |
Filaments pilose |
|
72 D. gyalanum |
|
|
|
|
164 (107) |
Leaves all basal. |
|
(165) |
+ |
Leaves both basal and cauline, sometimes basal and lower cauline leaves wilted at anthesis (in some alpine plants, lower internodes are very short and leaves may falsely appear to be all basal). |
|
(170) |
|
|
|
|
165 (164) |
Sepals adaxially puberulent. |
|
(166) |
+ |
Sepals adaxially glabrous. |
|
(167) |
|
|
|
|
166 (165) |
Lateral sepals more than 1.7 cm |
|
148 D. chayuense |
+ |
Lateral sepals ca. 1.7 cm |
|
143 D. thibeticum |
|
|
|
|
167 (165) |
Stem sparsely spreading pubescent; bracteoles more than 1 mm from flowers. |
|
(168) |
+ |
Stem retrorsely puberulent; bracteoles ca. 1 mm from flowers. |
|
(169) |
|
|
|
|
168 (167) |
Staminode limb suborbicular, entire; sepals elliptic-obovate or oblong, 1.5--1.8 cm |
|
125 D. autumnale |
+ |
Staminode limb broadly ovate, 2-cleft; sepals lanceolate, ca. 2.5 cm |
|
165 D. saxatile |
|
|
|
|
169 (167) |
Spur pendulous; sepals villous abaxially |
|
147 D. pomeense |
+ |
Spur horizontal; sepals puberulent abaxially |
|
149 D. sinoscaposum |
|
|
|
|
170 (164) |
Leaf blade primary lobes lobed for ca. 50% of their length, ultimate lobules ovate or narrowly so. |
|
(171) |
+ |
Leaf blade primary lobes lobed for more than 50% of their length, ultimate lobules lanceolate, narrowly so, or linear. |
|
(190) |
|
|
|
|
171 (170) |
Inflorescence less than 3 × as long as wide, or 1-flowered. |
|
(172) |
+ |
Inflorescence or branches ca. 3 × as long as wide. |
|
(176) |
|
|
|
|
172 (171) |
Leaf blade 1.3--2.6 cm wide. |
|
(173) |
+ |
Leaf blade 2.8--7 cm wide. |
|
(174) |
|
|
|
|
173 (172) |
Stem subglabrous; bracteoles leaflike, palmately 3-sect |
|
35 D. candelabrum |
+ |
Stem hispid; bracteoles linear, undivided |
|
145 D. pseudothibeticum |
|
|
|
|
174 (172) |
Pedicels 3.5--10 cm |
|
164 D. davidii |
+ |
Pedicels 1--2 cm. |
|
(175) |
|
|
|
|
175 (174) |
Spur ca. 2 cm; filaments glabrous |
|
38 D. henryi |
+ |
Spur ca. 1.5 cm; filaments sparsely hairy |
|
109 D. conaense |
|
|
|
|
176 (171) |
Staminode limb entire or emarginate at apex. |
|
(177) |
+ |
Staminode limb 2-lobed to 2-parted. |
|
(179) |
|
|
|
|
177 (176) |
Bracteoles ca. 1 mm from flowers, narrowly lanceolate; raceme with yellow, glandular hairs |
|
143 D. thibeticum |
+ |
Bracteoles more than 1 mm from flowers, narrowly linear or subulate; raceme without glandular hairs. |
|
(178) |
|
|
|
|
178 (177) |
Raceme rachis and pedicels densely retrorsely puberulent |
|
155 D. majus |
+ |
Raceme rachis glabrous, pedicels puberulent only at apex |
|
156 D. obcordatilimbum |
|
|
|
|
179 (176) |
Bracteoles ca. 1 mm from flowers. |
|
(180) |
+ |
Bracteoles more than 1 mm from flowers. |
|
(184) |
|
|
|
|
180 (179) |
Staminode limb with white hairs; leaf blade central segment undivided or indistinctly 3-lobed |
|
114 D. trifoliolatum |
+ |
Staminode limb with yellow or yellowish hairs; leaf blade central segment distinctly 3-lobed or 3-cleft. |
|
(181) |
|
|
|
|
181 (180) |
Stem ca. 10-leaved; basal leaves wilted at anthesis; carpels glabrous |
|
115 D. wangii |
+ |
Stem 1--3-leaved; basal leaves green at anthesis; carpels hairy. |
|
(182) |
|
|
|
|
182 (181) |
Stem spreading hispid |
|
127 D. yongningense |
+ |
Stem retrorsely pubescent or puberulent. |
|
(183) |
|
|
|
|
183 (182) |
Leaf blade 4.5--6 cm; raceme 7--10-flowered; filaments with a few hairs |
|
142 D. medogense |
+ |
Leaf blade 2.5--4 cm; raceme 2--4-flowered; filaments glabrous |
|
146 D. taipaicum |
|
|
|
|
184 (179) |
Pedicels glabrous. |
|
(185) |
+ |
Pedicels hairy. |
|
(187) |
|
|
|
|
185 (184) |
Stem hispid |
|
108 D. rangtangense |
+ |
Stem glabrous. |
|
(186) |
|
|
|
|
186 (185) |
Spur 1.8--2 cm, longer than sepals |
|
110 D. giraldii |
+ |
Spur 0.6--1.1 cm, subequaling sepals |
|
111 D. sparsiflorum |
|
|
|
|
187 (184) |
Pedicels with only yellow, glandular hairs |
|
107 D. pseudocampylocentrum |
+ |
Pedicels with white hairs (yellow, glandular hairs present or not). |
|
(188) |
|
|
|
|
188 (187) |
Inflorescence glabrous; sepals 0.7--0.8 cm |
|
112 D. angustipaniculatum |
+ |
Inflorescence hairy; sepals (0.9--)1.1--1.8 cm. |
|
(189) |
|
|
|
|
189 (188) |
Spur subulate-cylindric, 0.9--1 cm |
|
71 D. caudatolobum |
+ |
Spur subulate, 1.3--2.2 cm |
|
104 D. muliense |
|
|
|
|
190 (170) |
Inflorescence less than 3 × as long as wide, or 1-flowered. |
|
(191) |
+ |
Inflorescence ca. 3 × as long as wide. |
|
(204) |
|
|
|
|
191 (190) |
Bracteoles elliptic or elliptic-ovate; calyx urceolate; sepals longer than spur |
|
152 D. likiangense |
+ |
Bracteoles linear or subulate; calyx open; sepals shorter than spur. |
|
(192) |
|
|
|
|
192 (191) |
Bracteoles ca. 1 mm from flowers |
|
147 D. pomeense |
+ |
Bracteoles more than 1 mm from flowers. |
|
(193) |
|
|
|
|
193 (192) |
Staminode limb 2-cleft to middle. |
|
(194) |
+ |
Staminode limb emarginate or 2-lobulate at apex. |
|
(198) |
|
|
|
|
194 (193) |
Stem long branched proximally |
|
171 D. maximowiczii |
+ |
Stem simple or shortly branched distally. |
|
(195) |
|
|
|
|
195 (194) |
Ultimate leaf lobules narrowly ovate; spur 1.8--3 cm |
|
35 D. candelabrum |
+ |
Ultimate leaf lobules narrowly linear; spur ca. 2.5 cm. |
|
(196) |
|
|
|
|
196 (195) |
Stem simple; inflorescence 1-flowered; lateral sepals subequaling spur |
|
150 D. nortonii |
+ |
Stem branched distally; inflorescence 2--5-flowered; lateral sepals shorter than spur. |
|
(197) |
|
|
|
|
197 (196) |
Lateral sepals at least 1.7 cm; bracteoles borne proximally on pedicel, or absent |
|
164 D. davidii |
+ |
Lateral sepals less than 1.7 cm; bracteoles borne near middle of pedicel or distally |
|
170 D. pseudocaeruleum |
|
|
|
|
198 (193) |
Pedicels with spreading, white hairs (yellow, glandular hairs present or not). |
|
(199) |
+ |
Pedicels with retrorse or appressed, white hairs, or glabrous (yellow, glandular hairs present or not). |
|
(201) |
|
|
|
|
199 (198) |
Pedicels white puberulent only |
|
169 D. mollipilum |
+ |
Pedicels white puberulent and yellow glandular puberulent. |
|
(200) |
|
|
|
|
200 (199) |
Spur 2.5--3.1 cm; lateral sepals 2.4--3.6 cm; staminode limb ca. 13 × 10 mm |
|
167 D. grandilimbum |
+ |
Spur 3.3--3.8 cm; lateral sepals 1.4--1.6 cm; staminode limb ca. 6 × 5 mm |
|
168 D. pumilum |
|
|
|
|
201 (198) |
Stem proximally branched. |
|
(202) |
+ |
Stem simple or branched distally. |
|
(203) |
|
|
|
|
202 (201) |
Stem base retrorsely puberulent |
|
171 D. maximowiczii |
+ |
Stem base glabrous |
|
134 D. tenii |
|
|
|
|
203 (201) |
Sepals slightly shorter than spur; pedicels bracteolate above middle |
|
159 D. grandiflorum |
+ |
Sepals distinctly shorter than spur; pedicels bracteolate near middle |
|
166 D. tatsienense |
|
|
|
|
204 (190) |
Lateral sepals longer than spur. |
|
(205) |
+ |
Lateral sepals shorter than or equaling spur. |
|
(208) |
|
|
|
|
205 (204) |
Bracteoles ca. 1 mm from flowers, elliptic or elliptic-ovate; calyx urceolate; sepals 2--2.5 cm |
|
152 D. likiangense |
+ |
Bracteoles more than 1 mm from flowers, subulate; calyx open; sepals 1.1--1.8 cm. |
|
(206) |
|
|
|
|
206 (205) |
Raceme puberulent; staminode limb broadly obovate |
|
160 D. kamaonense |
+ |
Raceme glabrous; staminode limb obovate-oblong. |
|
(207) |
|
|
|
|
207 (206) |
Spur 1.3--1.5 cm; raceme 18--30-flowered |
|
137 D. pseudoyunnanense |
+ |
Spur 1.8--2 cm; raceme 5--9-flowered |
|
141 D. pycnocentrum |
|
|
|
|
208 (204) |
Spur subequaling lateral sepals, 1--1.4 cm, abruptly upcurved |
|
138 D. yuanum |
+ |
Spur usually longer than lateral sepals, (1.2--)1.5--3 cm, horizontal or recurved, rarely slightly upcurved. |
|
(209) |
|
|
|
|
209 (208) |
Staminode limb glabrous; sepals ± forward pointing |
|
151 D. souliei |
+ |
Staminode limb barbate; sepals ± spreading. |
|
(210) |
|
|
|
|
210 (209) |
Bracteoles ca. 1 mm from flower. |
|
(211) |
+ |
Bracteoles more than 1 mm from flowers. |
|
(214) |
|
|
|
|
211 (210) |
Raceme dense, pedicels densely yellow glandular puberulent |
|
116 D. aemulans |
+ |
Raceme ± sparse, pedicels only white puberulent, rarely mixed with a few yellow, glandular hairs. |
|
(212) |
|
|
|
|
212 (211) |
Cauline leaf blade ultimate lobules pungent at apex |
|
143 D. thibeticum |
+ |
Cauline leaf blade ultimate lobules acute or slightly obtuse at apex. |
|
(213) |
|
|
|
|
213 (212) |
Raceme at least 5-flowered; bracteoles lanceolate, 1--1.6 mm wide; spur 1.9--2.7 cm |
|
143 D. thibeticum |
+ |
Raceme 2--4-flowered; bracteoles oblong-ovate, 3--3.5 mm wide; spur 1.4--2 cm |
|
146 D. taipaicum |
|
|
|
|
214 (210) |
Raceme glabrous. |
|
(215) |
+ |
Raceme ± hairy. |
|
(218) |
|
|
|
|
215 (214) |
Carpels glabrous |
|
135 D. shuichengense |
+ |
Carpels hairy. |
|
(216) |
|
|
|
|
216 (215) |
Stem glabrous |
|
156 D. obcordatilimbum |
+ |
Stem hairy proximally, glabrous distally. |
|
(217) |
|
|
|
|
217 (216) |
Pedicels yellow pubescent near apex |
|
125 D. autumnale |
+ |
Pedicels glabrous |
|
132 D. leiostachyum |
|
|
|
|
218 (214) |
Raceme sparsely puberulent. |
|
(219) |
+ |
Raceme densely puberulent. |
|
(222) |
|
|
|
|
219 (218) |
Staminode limb suborbicular, apex entire |
|
125 D. autumnale |
+ |
Staminode limb obovate, apex usually emarginate or 2-lobed, rarely entire. |
|
(220) |
|
|
|
|
220 (219) |
Raceme dense; ovary sparsely pubescent |
|
141 D. pycnocentrum |
+ |
Raceme lax; ovary densely pubescent. |
|
(221) |
|
|
|
|
221 (220) |
Stem more than 50 cm |
|
136 D. yunnanense |
+ |
Stem to 50 cm |
|
160 D. kamaonense |
|
|
|
|
222 (218) |
Staminode limb broadly obovate or suborbicular |
|
159 D. grandiflorum |
+ |
Staminode limb narrowly obovate or oblong. |
|
(223) |
|
|
|
|
223 (222) |
Bracteoles narrowly oblong, 11--18 × 3--4 mm |
|
157 D. weiningense |
+ |
Bracteoles linear or subulate, 2.5--7 × 0.2--1 mm. |
|
(224) |
|
|
|
|
224 (223) |
Sepals abaxially only puberulent, spur 1.6--2.2 cm; petals glabrous; staminode limb emarginate at apex |
|
155 D. majus |
+ |
Sepals abaxially puberulent and villous, spur 2.4--2.8 cm; petals ciliate at apex; staminode limb erose at apex |
|
158 D. nangchienense |