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87. Deyeuxia Clarion ex P. Beauvois, Ess. Agrostogr. 43. 1812.

野青茅属 ye qing mao shu

Authors: Sheng-lian Lu, Wenli Chen & Sylvia M. Phillips

Anisachne Keng.

Perennials, tufted or also rhizomatous. Leaf blades linear, usually flat, sometimes inrolled; ligule membranous. Inflorescence a panicle, loosely contracted to spikelike, rarely open. Spikelets with 1 floret (very rarely 2), rachilla disarticulating above glumes, with penicillate extension lying against the palea; glumes subequal, usually slightly longer than the floret, rarely slightly shorter, membranous, 1–3-veined, apex acute or acuminate; floret callus bearded, hairs usually 1/3 as long up to about equaling floret; lemma thinly to firmly membranous, (3–)5-veined, dorsally awned or awnless, apex erose, denticulate or 2–4-toothed; awn geniculate or straight, its position varying from near base to near apex, usually inconspicuous, up to twice length of lemma but occasionally reduced to a subapical mucro or absent; palea 2/3 as long to subequaling lemma. Stamens usually 3, rarely 2 or 1.

About 200 species:temperate regions throughout the world, also on tropical mountains; 34 species (15 endemic) in China.

The genera Agrostis, Calamagrostis, and Deyeuxia form an intergrading complex of three incompletely separated entities. The majority of species can be placed without difficulty, but there is a number of troublesome intermediates, and no character combinations provide a definitive way of distinguishing the genera. Agrostis, with small spikelets and short callus hairs, has always been maintained separately, while Deyeuxia is sometimes maintained and sometimes sunk into Calamagrostis. However, the boundary between Agrostis and Calamagrostis is just as ill-defined as that between Calamagrostis and Deyeuxia. For this reason, three separate genera are maintained here, which complies with the usual practice in Chinese floras and avoids the need for many new combinations.

The taxonomy of this group is complicated by the presence of a number of apomictic complexes and frequent hybridization. Many infraspecific taxa have been described in an attempt to order this variation. However, intermediates are to be expected in an actively evolving group such as this, and most of these infraspecific taxa have not been enumerated separately here.


1 Lemma awnless; panicle open   (2)
+ Lemma awned; panicle open or contracted   (6)
       
2 (1) Callus hairs 2/3 as long as lemma or more; culms up to 120 cm; panicle 15–30 cm   (3)
+ Callus hairs not more than 1/2 as long as lemma; culms up to 60 cm; panicle 6–12 cm   (4)
       
3 (2) Spikelets 2–3 mm; lemma 1.5–2.5 mm; callus hairs equaling or subequaling lemma.   1 D. diffusa
+ Spikelets 3–4.2 mm; lemma 3–4 mm; callus hairs 2/3–4/5 as long as lemma.   2 D. flaccida
       
4 (2) Spikelets 3–3.7 mm; rachilla 1.5–2.5 mm; anthers ca. 2 mm.   3 D. yanyuanensis
+ Spikelets 2–2.5 mm; rachilla ca. 0.6 mm; anthers ca. 0.5 mm   (5)
       
5 (4) Spikelets 1.9–2.5(–3) mm; glumes slightly shorter than floret; callus hairs less than 1 mm, not visible between glumes; leaf sheaths smooth.   4 D. petelotii
+ Spikelets 2.5–3.5 mm; glumes equaling or longer than floret; callus hairs 1.2–2 mm, visible between glumes; leaf sheaths scabrid   5 D. abnormis
       
6 (1) Callus hairs less than 1/2 as long as lemma   (7)
+ Callus hairs 1/2 as long as lemma or longer   (19)
       
7 (6) Awn arising near or above middle of lemma   (8)
+ Awn arising from below middle of lemma   (12)
       
8 (7) Spikelets with 2 florets.   6 D. himalaica
+ Spikelets with 1 floret   (9)
       
9 (8) Plant with slender, elongate, creeping rhizomes   (10)
+ Plant tufted or shortly rhizomatous   (11)
       
10 (9) Awn 1–2.5(–6) mm, arising from upper 1/3 of lemma or above; glumes scabrid.   7 D. pulchella
+ Awn 5–6 mm, arising from middle of lemma; glumes smooth or scabrid only on keel.   8 D. rosea
       
11 (9) Panicle contracted, branches straight, scabrid; lower glume ciliolate on margins; anthers 2–3 mm.   9 D. scabrescens
+ Panicle open, branches capillary, curving, smooth; lower glume not ciliolate on margins; anthers ca. 1 mm.   10 D. nyingchiensis
       
12 (7) Panicle loosely contracted to dense, sometimes spikelike   (13)
+ Panicle open with spreading branches, (5–)15–35 cm wide   (16)
       
13 (12) Anthers 0.5–1.2 mm; culms 1–2-noded.   11 D. nivicola
+ Anthers 2–3 mm; culms (2–)3–5-noded   (14)
       
14 (13) Glumes membranous, shining.   25 D. anthoxanthoides
+ Glumes not shining   (15)
       
15 (14) Leaf blades involute, filiform; spikelets 4–5.5 mm.   12 D. mazzettii
+ Leaf blades flat, stiff; spikelets 5–7 mm.   14 D. korotkyi
       
16 (12) Callus hairs less than 1/5 length of lemma; ligule 0.5–2(–4) mm   (17)
+ Callus hairs 1/5–2/5 length of lemma; ligule 2.5–20 mm   (18)
       
17 (16) Plant ca. 65 cm tall; panicle 5–9(–16) cm; spikelets with 1(–2) florets.   15 D. suizanensis
+ Plant 80–120 cm tall; panicle 20–35 cm; spikelets with 1 floret.   16 D. effusiflora
       
18 (16) Lemma apex erose; anthers (1.3–)2–3 mm.   17 D. pyramidalis
+ Lemma apex with 0.5–1.5 mm mucros; anthers 0.5–1.2 mm.   13 D. flavens
       
19 (6) Panicle open or slightly contracted   (20)
+ Panicle dense, often spikelike   (24)
       
20 (19) Panicle branches usually paired; glumes smooth, scabrid on keel only; ligule 1–1.5 mm.   18 D. hakonensis
+ Panicle branches fascicled; glumes scabrid; ligule 3–20 mm   (21)
       
21 (20) Awn sub-basal.   19 D. sinelatior
+ Awn arising near or above middle of lemma   (22)
       
22 (21) Culms (4–)6–8-noded, usually branched.   20 D. purpurea
+ Culms 2–4-noded, unbranched   (23)
       
23 (22) Lemma awned from near middle; awn ca. 1.2 mm, not exserted from spikelets.   21 D. sichuanensis
+ Lemma awned from near apex; awn 3–6 mm, exserted from spikelet.   22 D. nepalensis
       
24 (19) Awn geniculate with twisted column, conspicuously exceeding glumes   (25)
+ Awn slender, straight, slightly curved or indistinctly twisted in lower part, included within glumes (exserted in D. moupinensis)   (27)
       
25 (24) Culm densely pubescent below panicle; panicle branches pubescent.   26 D. tibetica
+ Culms scabrid below panicle; panicle branches scabrid   (26)
       
26 (25) Anthers 1.5–2 mm; spikelets 4–6 mm, purple.   27 D. tianschanica
+ Anthers 2–4 mm; spikelets 5–9 mm, yellowish brown, purplish at base.   28 D. holciformis
       
27 (24) Lemma apex deeply 2-lobed; awn arising from between lobes; ligule ca. 0.5 mm.   29 D. moupinensis
+ Lemma apex denticulate or minutely 4-toothed; ligule 1.5–7 mm   (28)
       
28 (27) Anthers ca. 0.5 mm; culms 15–45 cm tall.   30 D. debilis
+ Anthers 1.5–2.2 mm   (29)
       
29 (28) Awn arising from lower 1/3 of lemma or slightly above   (30)
+ Awn sub-basal   (32)
       
30 (29) Awn poorly developed, 1–2 mm; spikelets 3–4 mm.   23 D. neglecta
+ Awn 3–6 mm; spikelets 4–7 mm   (31)
       
31 (30) Panicle 10–25 cm; glumes subequal.   24 D. lapponica
+ Panicle 5–8 cm; glumes unequal.   31 D. zangxiensis
       
32 (29) Callus hairs equal to or slightly longer than lemma.   32 D. conferta
+ Callus hairs 1/2–4/5 as long as lemma   (33)
       
33 (32) Leaf sheaths scabrid; spikelets purple with bronze at apex.   33 D. kokonorica
+ Leaf sheaths smooth; spikelets yellowish green or pinkish.   34 D. macilenta

  • List of lower taxa


     

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